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	<dc:identifier>FB-9563</dc:identifier>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=9563</dc:source>
	<dwc:Kingdom>Animalia</dwc:Kingdom>
	<dwc:Phylum>Chordata</dwc:Phylum>
	<dwc:Class>Teleostei</dwc:Class>
	<dwc:Order>Siluriformes</dwc:Order>
	<dwc:Family>Mochokidae</dwc:Family>
	<dwc:Genus>Synodontis</dwc:Genus>
	<dwc:ScientificName>Synodontis frontosus Vaillant, 1895</dwc:ScientificName>
	<commonName xml:lang=''>Kwoke</commonName>
	<commonName xml:lang='ar'>Gargur</commonName>
	<commonName xml:lang='en'>Sudan squeaker</commonName>
	<commonName xml:lang='fi'>Sudaninripsimonni</commonName>
	<commonName xml:lang=''>Gbagagbaga</commonName>
	<commonName xml:lang=''>Lour kasicou</commonName>
	<commonName xml:lang=''>Pua</commonName>
	<commonName xml:lang=''>大額歧鬚鮠</commonName>
	<commonName xml:lang=''>大额歧须鮠</commonName>
	<synonym relationship='synonym'>Synodontis citernii Vinciguerra, 1898</synonym>
	<synonym relationship='synonym'>Synodontis frontosa Vaillant, 1895</synonym>
	<dcterms:created>1994-04-21</dcterms:created>
	<dcterms:modified>2024-08-12</dcterms:modified>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=205&amp;speccode=9563'>Breder, C.M. and D.E. Rosen0 Modes of reproduction in fishes. T.F.H. Publications, Neptune City, New Jersey. 941 p. (Ref. 205)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=367&amp;speccode=9563'>Blache, J.0 Les poissons du bassin du Tchad et du bassin adjacent du Mayo Kebbi. Étude systématique et biologique. Mém. O.R.S.T.O.M. 4(2):483 p. (Ref. 367)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=3202&amp;speccode=9563'>Gosse, J.-P.0 Mochokidae. p. 105-152. In J. Daget, J.-P. Gosse and D.F.E. Thys van den Audenaerde (eds.) Check-list of the freshwater fishes of Africa (CLOFFA). ISNB, Brussels, MRAC, Tervuren; and ORSTOM, Paris. Vol. 2. (Ref. 3202)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4517&amp;speccode=9563'>Hureau, J.-C.0 La base de données GICIM: Gestion informatisée des collections ichthyologiques du Muséum. p. 225-227. In Atlas Préliminaire des Poissons d'Eau Douce de France. Conseil Supérieur de la Pêche, Ministère de l'Environnement, CEMAGREF et Muséum national d'Histoire naturelle, Paris. (Ref. 4517)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4903&amp;speccode=9563'>Greenwood, P.H.0 The Fishes of Uganda. The Uganda Society, Kampala. 131p. (Ref. 4903)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4906&amp;speccode=9563'>Bianco, P.0 Fish. p. 55-63. In L. Boitani (ed.) The Southern National Park. A master plan. Istituto di Zoologia, Facoltà di Scienze dell'Università di Roma (Italy). 220 p. (Ref. 4906)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=6160&amp;speccode=9563'>Hickley, P. and R.G. Bailey0 Food and feeding relationships of fish in the Sudd swamps (River Nile, southern Sudan). J. Fish Biol. 30(2):147-159. (Ref. 6160)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=7362&amp;speccode=9563'>Paugy, D. and T.R. Roberts0 Mochokidae. p. 500-563. In C. Levêque, D. Paugy, and G.G. Teugels (eds.) Faune des poissons d'eaux douces et saumâtres d'Afrique de l'Ouest Tome 2. Coll. Faune Tropicale n° 28. Musée Royal de l'Afrique Centrale, Tervuren, Belgique and O.R.S.T.O.M., Paris, France, 902. (Ref. 7362)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=12818&amp;speccode=9563'>Anonymous0 Fish registrations within the museum database of the Vertebrate Section of the Royal Museum for Central Africa. MRAC, Tervuren, Belgium. (Ref. 12818)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=28714&amp;speccode=9563'>Bailey, R.G.0 Guide to the fishes of the River Nile in the Republic of the Sudan. J. Nat. Hist. 28:937-970. (Ref. 28714)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=31439&amp;speccode=9563'>Boulenger, G.A.0 Zoology of Egypt: the fishes of the Nile. Hugh Rees, Limited, London. 578p. (Ref. 31439)</reference>
	<reference isbn='957-8596-59-6' url='http://www.fishbase.org/references/FBRefSummary.php?id=31517&amp;speccode=9563'>Wu, H.L., K.-T. Shao and C.F. Lai (eds.)0 Latin-Chinese dictionary of fishes names. The Sueichan Press, Taiwan. 1028 p. (Ref. 31517)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=31982&amp;speccode=9563'>Anonymous0 Fish collection database of the Natural History Museum, London (formerly British Museum of Natural History (BMNH)). Natural History Museum, London (formerly British Museum of Natural History (BMNH)). (Ref. 31982)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=35508&amp;speccode=9563'>Anonymous0 The icthyological collection of the Zoological Museum Hamburg (ZMH). Division of Ichthyology and Herpetology, Zoological Museum Hamburg (ZMH). (Ref. 35508)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=36900&amp;speccode=9563'>Copley, H.0 Common freshwater fishes of East Africa. H.F. &amp; G. Witherby Ltd., London. 172p. (Ref. 36900)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=38732&amp;speccode=9563'>Anonymous0 Fish collection database of the National Museum of Natural History (Smithsonian Institution). Smithsonian Institution-Division of Fishes. (Ref. 38732)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=41414&amp;speccode=9563'>Anonymous0 Fish collection database of the American Museum of Natural History. American Museum of Natural History, Central Park West , NY 10024-5192, USA. (Ref. 41414)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=43507&amp;speccode=9563'>Worthington, E.B. and C.K. Ricardo0 Scientific results of the Cambridge Expedition to the east African lakes, 1930-1. No. 15. The fish of Lake Rudolf and Lake Baringo. J. Linn. Soc. (Zool.) 39(267):353-389. (Ref. 43507)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=49805&amp;speccode=9563'>Worthington, E.B.0 A report on the fishing survey of lakes Albert and Kioga. March to July, 1928. Crown Agents for the Colonies, London. 136 p. (Ref. 49805)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=51086&amp;speccode=9563'>Varjo, M., L. Koli and H. Dahlström0 Kalannimiluettelo (versio 10/03). Suomen Biologian Seura Vanamo Ry. (Ref. 51086)</reference>
	<reference isbn='2.11.086730-2' url='http://www.fishbase.org/references/FBRefSummary.php?id=51193&amp;speccode=9563'>Vivien, J.0 Faune du Cameroun. Guide des mammifères et poissons. GICAM et Ministère de la Coopération et du Développement, Yaoundé/Paris. 271 p. (Ref. 51193)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=52331&amp;speccode=9563'>Seegers, L., L. De Vos and D.O. Okeyo0 Annotated checklist of the freshwater fishes of Kenya (excluding the lacustrine haplochromines from Lake Victoria). J. E. Afr. Nat. Hist. 92:11-47. (Ref. 52331)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=52871&amp;speccode=9563'>Okeyo, D.O.0 Taxonomy, common names and distribution of fish in the eastern arm of the Rift Valley drainage, Kenya. p. 250-266. In M.L.D. Palomares, B. Samb, T. Diouf, J.M. Vakily and D. Pauly (eds.) Fish biodiversity: local studies as basis for global inferences. ACP-EU Fish. Res. Rep. 14:281 p. (Ref. 52871)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57223&amp;speccode=9563'>Paugy, D. and T.R. Roberts0 Mochokidae. p. 195-268 In C. Lévêque, D. Paugy and G.G. Teugels (eds.) Faune des poissons d'eaux douce et saumâtres de l'Afrique de l'Ouest, Tome 2. Coll. Faune et Flore tropicales 40. Musée Royal de l'Afrique Centrale, Tervuren, Belgique, Museum National d'Histoire Naturalle, Paris, France and Institut de Recherche pour le Développement, Paris, France. 815 p. (Ref. 57223)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=58032&amp;speccode=9563'>Ferraris, C.J. Jr.0 Checklist of catfishes, recent and fossil (Osteichthyes: Siluriformes), and catalogue of siluriform primary types. Zootaxa 1418:1-628. (Ref. 58032)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=58108&amp;speccode=9563'>Chinese Academy of Fishery Sciences0 Chinese aquatic germplasm resources database. http://zzzy.fishinfo.cn (Ref. 58108)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=58460&amp;speccode=9563'>Getahun, A.0 An overview of the diversity and conservation status of the Ethiopian freshwater fauna. J. Afrotrop. Zool. Special Issue:87-96. (Ref. 58460)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=82238&amp;speccode=9563'>Poll, M.0 Révision des &lt;i&gt;Synodontis&lt;/i&gt; africains (famille Mochocidae). Ann. Mus. R. Afr. Centr., Sci. Zool., 191:1-497. (Ref. 82238)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=87720&amp;speccode=9563'>Greenwood, P.H.0 The fishes of Uganda - III. Uganda J. 21(1):47-80. (Ref. 87720)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=92382&amp;speccode=9563'>Vaillant, L.0 Sur la dentition des &lt;i&gt;Synodontis&lt;/i&gt; et diagnose d'espèces nouvelles. C. R. Soc. Philomat. Paris 16:47-48. (Ref. 92382)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=97924&amp;speccode=9563'>Nyingi, W.D.0 Guide to common freshwater fishes of Kenya. Moran Publishers, Nairobi, Kenya. 136 p. (Ref. 97924)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=120561&amp;speccode=9563'>Moritz, T., Z.N. El Dayem, M.A. Abdallah and D. Neumann0 New and rare records of fishes from the White Nile in the Republic of the Sudan. Cybium 43(2):137-151. (Ref. 120561)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=137577&amp;speccode=9563'>Price, S.A., S.T. Friedman, K.A. Corn, O. Larouche, K. Brockelsby, A.J. Lee, M. Nagaraj, N.G. Bertrand, M. Danao, M.C. Coyne, J.R. Estrada, R. Friedman, E. Hoeft, M. Iwan, D. Gross, J.H. Kao, B. Landry, M.J. Linares, C. McGlinn and …0 FishShapes v1: Functionally relevant measurements of teleost shape and size on three dimensions. Ecology 103(e3829):1-2. (Ref. 137577)</reference><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Pascualita Sa-a</agent>
	<dcterms:created>1994-04-21 00:00:00</dcterms:created>
	<dcterms:modified>2024-08-12 00:00:00</dcterms:modified>
	<dc:title>Biology</dc:title>
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	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=9563</dc:source>
	<dc:description>Found in deep, open waters and shallow inshore areas of Lake Albert (Ref. 49805). Occurs in lakes and channels; feeds on algae, macrophytes, detritus, crustaceans, insects and mollusks; breeding takes place during flooding in June to early dry down in January  (Ref. 28714). Oviparous (Ref. 205).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57223&amp;speccode=9563'>Paugy, D. and T.R. Roberts 2003 Mochokidae. p. 195-268 In C. Lévêque, D. Paugy and G.G. Teugels (eds.) Faune des poissons d'eaux douce et saumâtres de l'Afrique de l'Ouest, Tome 2. Coll. Faune et Flore tropicales 40. Musée Royal de l'Afrique Centrale, Tervuren, Belgique, Museum National d'Histoire Naturalle, Paris, France and Institut de Recherche pour le Développement, Paris, France. 815 p. (Ref. 57223)</reference></dataObject><dataObject>
	<dc:identifier>FB-DiagnosticDescr-9563</dc:identifier>
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	<agent homepage='http://www.africamuseum.be#' logoURL='' role='recorder'>Gert Boden</agent>
	<dcterms:created>2012-12-13 00:00:00</dcterms:created>
	<dcterms:modified>2015-03-17 21:24:32</dcterms:modified>
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	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
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	<audience>General public</audience>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Size</subject>
	<dc:description>Diagnosis: gill slits not extending ventrally beyond pectoral-fin insertions; interorbital space over 50% of head width; post-orbital length at least 60% of snout length; maxillary barbels longer than head, unbranched, without tubercles and bordered by a short but distinct, dark basal membrane; outer mandibular barbels with rather numerous, simple ramifications, branches of inner mandibular barbels subdivided, thick and tuberculate; mandibular teeth rather short, numbering 36-48; denticulations of pectoral-fin spines strong on inner, and hardly visible on outer margin; dorsal-fin spine smooth anteriorly, except for some apical denticles; first branched ray of dorsal and pectoral fins prolonged into a filament; adipose fin high and long, distincly separated from rayed dorsal fin; humeral process deep, pointed and granulose, not keeled ventrally; body covered with very small black specks; caudal fin without spots or dots (Ref. 57223).Coloration: uniformly brownish, with numerous very small black spots, also on adipose fin, but none on other fins (Ref. 57223).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57223&amp;speccode=9563'>Paugy, D. and T.R. Roberts 2003 Mochokidae. p. 195-268 In C. Lévêque, D. Paugy and G.G. Teugels (eds.) Faune des poissons d'eaux douce et saumâtres de l'Afrique de l'Ouest, Tome 2. Coll. Faune et Flore tropicales 40. Musée Royal de l'Afrique Centrale, Tervuren, Belgique, Museum National d'Histoire Naturalle, Paris, France and Institut de Recherche pour le Développement, Paris, France. 815 p. (Ref. 57223)</reference></dataObject><dataObject>
	<dc:identifier>FB-Morphology-9563</dc:identifier>
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	<agent homepage='http://www.africamuseum.be#' logoURL='' role='recorder'>Gert Boden</agent>
	<dcterms:created>2012-12-13 00:00:00</dcterms:created>
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	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
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	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=9563</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Threats</subject>
	<dc:description>Dorsal spines (total): 1; Dorsal soft rays (total): 7;  Analspines: 0; Analsoft rays: 13</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57223&amp;speccode=9563'>Paugy, D. and T.R. Roberts 2003 Mochokidae. p. 195-268 In C. Lévêque, D. Paugy and G.G. Teugels (eds.) Faune des poissons d'eaux douce et saumâtres de l'Afrique de l'Ouest, Tome 2. Coll. Faune et Flore tropicales 40. Musée Royal de l'Afrique Centrale, Tervuren, Belgique, Museum National d'Histoire Naturalle, Paris, France and Institut de Recherche pour le Développement, Paris, France. 815 p. (Ref. 57223)</reference></dataObject><dataObject>
	<dc:identifier>FB-Distribution-9563</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Pascualita Sa-a</agent>
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	<dcterms:rightsHolder>WorldFish Center - FishBase</dcterms:rightsHolder>
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	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=9563</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Habitat</subject>
	<dc:description>Africa:  Nile basin, LakesTurkana, Albert, Edward and Kioga (Ref. 3202, 82238), and Shabeelle River (Ref. 3202, 82238). Reported from Lake Edward (Ref. 3202, 82238), but Mochokidae are not present in the lake (Ref. 120232). Its presence in Chad basin (Ref. 367) is questionable and needs confirmation (Ref. 7362, 57223, 82238).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57223&amp;speccode=9563'>Paugy, D. and T.R. Roberts 2003 Mochokidae. p. 195-268 In C. Lévêque, D. Paugy and G.G. Teugels (eds.) Faune des poissons d'eaux douce et saumâtres de l'Afrique de l'Ouest, Tome 2. Coll. Faune et Flore tropicales 40. Musée Royal de l'Afrique Centrale, Tervuren, Belgique, Museum National d'Histoire Naturalle, Paris, France and Institut de Recherche pour le Développement, Paris, France. 815 p. (Ref. 57223)</reference></dataObject><dataObject>
	<dc:identifier>FB-Uses-9563</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Pascualita Sa-a</agent>
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	<dc:title>Importance</dc:title>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution</subject>
	<dc:description>fisheries: of no interest</dc:description>
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	<dc:identifier>FB-LifeCycle-9563</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Susan M. Luna</agent>
	<dcterms:created>1994-04-21 00:00:00</dcterms:created>
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	<dc:source>http://www.fishbase.org//Reproduction/FishReproSummary.php?ID=9563&amp;StockCode=9886</dc:source>
	<dc:description>Oviparous (Ref. 205). Distinct pairing during breeding (Ref. 205).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=205&amp;speccode=9563'>Breder, C.M. and D.E. Rosen 1966 Modes of reproduction in fishes. T.F.H. Publications, Neptune City, New Jersey. 941 p. (Ref. 205)</reference></dataObject><dataObject>
	<dc:identifier>FB-TrophicStrategy-9563</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Pascualita Sa-a</agent>
	<dcterms:created>1997-03-14 00:00:00</dcterms:created>
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	<dc:source>frontosus</dc:source>
	<dc:description>Feeds on detritus, plants and insects (Ref. 6160).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=6160&amp;speccode=9563'>Hickley, P. and R.G. Bailey 1987 Food and feeding relationships of fish in the Sudd swamps (River Nile, southern Sudan). J. Fish Biol. 30(2):147-159. (Ref. 6160)</reference></dataObject><dataObject>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57223&amp;speccode=9563'>Paugy, D. and T.R. Roberts 2003 Mochokidae. p. 195-268 In C. Lévêque, D. Paugy and G.G. Teugels (eds.) Faune des poissons d'eaux douce et saumâtres de l'Afrique de l'Ouest, Tome 2. Coll. Faune et Flore tropicales 40. Musée Royal de l'Afrique Centrale, Tervuren, Belgique, Museum National d'Histoire Naturalle, Paris, France and Institut de Recherche pour le Développement, Paris, France. 815 p. (Ref. 57223)</reference></dataObject><dataObject>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57073'>IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006.</reference></dataObject><dataObject>
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