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<taxon>
	<dc:identifier>FB-2319</dc:identifier>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=2319</dc:source>
	<dwc:Kingdom>Animalia</dwc:Kingdom>
	<dwc:Phylum>Chordata</dwc:Phylum>
	<dwc:Class>Teleostei</dwc:Class>
	<dwc:Order>Siluriformes</dwc:Order>
	<dwc:Family>Clariidae</dwc:Family>
	<dwc:Genus>Heterobranchus</dwc:Genus>
	<dwc:ScientificName>Heterobranchus longifilis Valenciennes, 1840</dwc:ScientificName>
	<commonName xml:lang='af'>Vundu</commonName>
	<commonName xml:lang='ar'>Surta</commonName>
	<commonName xml:lang=''>Sampa</commonName>
	<commonName xml:lang=''>Vundu</commonName>
	<commonName xml:lang='da'>Vundu-malle</commonName>
	<commonName xml:lang='en'>Catfish</commonName>
	<commonName xml:lang='en'>Sampa</commonName>
	<commonName xml:lang='en'>Vundu</commonName>
	<commonName xml:lang='ee'>Wonyi</commonName>
	<commonName xml:lang='fi'>Oliiviahmattimonni</commonName>
	<commonName xml:lang='fi'>Viiksiankeriasmonni</commonName>
	<commonName xml:lang=''>Denor</commonName>
	<commonName xml:lang='ha'>Ramboshi</commonName>
	<commonName xml:lang='ig'>Azu isii</commonName>
	<commonName xml:lang='kr'>Kairo</commonName>
	<commonName xml:lang='sw'>Pondo</commonName>
	<commonName xml:lang='sw'>Vundu</commonName>
	<commonName xml:lang='wo'>Biblique</commonName>
	<commonName xml:lang='yo'>Aso</commonName>
	<commonName xml:lang=''>Coro</commonName>
	<commonName xml:lang=''>Cur</commonName>
	<commonName xml:lang=''>Fusan</commonName>
	<commonName xml:lang=''>Giyoko</commonName>
	<commonName xml:lang=''>Isu</commonName>
	<commonName xml:lang=''>Labe</commonName>
	<commonName xml:lang=''>Lenda</commonName>
	<commonName xml:lang=''>Lifo</commonName>
	<commonName xml:lang=''>Lifonde</commonName>
	<commonName xml:lang=''>Ngaru</commonName>
	<commonName xml:lang=''>Nhumi</commonName>
	<commonName xml:lang=''>Seega</commonName>
	<commonName xml:lang=''>Su</commonName>
	<commonName xml:lang=''>Sumi</commonName>
	<commonName xml:lang=''>Toruye</commonName>
	<commonName xml:lang=''>Tshota mualwa</commonName>
	<commonName xml:lang=''>長絲異鰓鯰</commonName>
	<commonName xml:lang=''>长丝异鳃鲶</commonName>
	<synonym relationship='synonym'>Clarias loangwensis Worthington, 1933</synonym>
	<synonym relationship='synonym'>Heterobranchus laticeps Peters, 1852</synonym>
	<synonym relationship='synonym'>Heterobranchus longifinis Valenciennes, 1840</synonym>
	<synonym relationship='synonym'>Heterobranchus platycephalus Nichols &amp; LaMonte, 1934</synonym>
	<dcterms:created>1991-02-27</dcterms:created>
	<dcterms:modified>2024-01-22</dcterms:modified>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=55876&amp;speccode=2319'>Adebisi, A.A.0 Analyses of the stomach contents of the piscivorous fishes of the upper Ogun River in Nigeria. Hydrobiologia 79(2):167-177. (Ref. 55876)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=56439&amp;speccode=2319'>Balon, E.K.0 Possible fish stock size assessment and available production survey as developed on Lake Kariba. Afr. J. Trop. Hydrobiol. Fish. 2:45-73. (Ref. 56439)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57129&amp;speccode=2319'>Teugels, G.G.0 Clariidae. p. 144-173 In C. Lévêque, D. Paugy and G.G. Teugels (eds.) Faune des poissons d'eaux douce et saumâtres de l'Afrique de l'Ouest, Tome 2. Coll. Faune et Flore tropicales 40. Musée Royal de l'Afrique Centrale, Tervuren, Belgique, Museum National d'Histoire Naturalle, Paris, France and Institut de Recherche pour le Développement, Paris, France. 815 p. (Ref. 57129)</reference>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=58460&amp;speccode=2319'>Getahun, A.0 An overview of the diversity and conservation status of the Ethiopian freshwater fauna. J. Afrotrop. Zool. Special Issue:87-96. (Ref. 58460)</reference>
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	<reference isbn='978-3-936027-83-9' url='http://www.fishbase.org/references/FBRefSummary.php?id=78218&amp;speccode=2319'>Seegers, L.0 The catfishes of Africa: A handbook for identification and maintenance. Aqualog Verlag A.C.S. GmbH, Germany. 604 p. (Ref. 78218)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=81208&amp;speccode=2319'>Coulibaly, A., T. Koné, N.I. Ouattara, V. N'Douba, J. Snoeks, E.P. Kouamélan and G. Gooré Bi0 Évaluation de l'effet d'un système de refuge sur la survie et la croissance des alevins de &lt;i&gt;Heterobranchus longifilis&lt;/i&gt; élevés en cage flottante. Belg. J. Zool. 137(2):157-164. (Ref. 81208)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=81644&amp;speccode=2319'>Teugels, G.G., D. Adriaens, S. Devaere and T. Musschoot0 Clariidae. p. 653-691. In M.L.J. Stiassny, G.G. Teugels and C.D. Hopkins (eds.) The fresh and brackish water fishes of Lower Guinea, West-Central Africa. Volume I. Collection Faune et Flore tropicales 42. Institut de Recherche pour le Développement, Paris, France, Muséum National d’Histoire Naturelle, Paris, France, and Musée Royal de l’Afrique Centrale, Tervuren, Belgium. 800 pp. (Ref. 81644)</reference>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=83625&amp;speccode=2319'>Konan, F.K., F. Leprieur, A. Ouattara, S. Brosse, G. Grenouillet, G. Gourène, P. Winterton and S. Lek0 Spatio-temporal patterns of fish assemblages in coastal West African rivers: a self-organizing map approach. Aquat. Living Resour. 19(4):361-370. (Ref. 83625)</reference>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=93897&amp;speccode=2319'>Monsembula Iyaba, R.J.C. and M.L.J. Stiassny0 Fishes of the Salonga National Park (Congo basin, central Africa): a list of species collected in the Luilaka, Salonga, and Yenge Rivers (Equateur Province, Democratic Republic of Congo). Check List 9(2):246-256. (Ref. 93897)</reference>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=102072&amp;speccode=2319'>Inyang, N.M., C.I.P. Anibeze and B.O. Mbbenka0 Aspects of the reproduction biology of &lt;i&gt;Heterobranchus longifilis&lt;/i&gt; (Valenciennes, 1840) in the Idodo River Basin, southeastern Nigeria. J. Afr. Zool. 111(5):373-379. (Ref. 102072)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=106245&amp;speccode=2319'>Moelants, T.0 Diversity and ecology of the ichthyofauna of the Middle and Upper Congo basin: a case-study in the region of the Wagenia falls (Democratic Republic of the Congo). KULeuven, Faculty of Science, Leuven (Belgium), 320p. (Ref. 106245)</reference>
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	<reference isbn='9781868107186' url='http://www.fishbase.org/references/FBRefSummary.php?id=115164&amp;speccode=2319'>Okeyo, D.O. and W.O. Ojwang (Comp.)0 A photographic guide to freshwater fishes of Kenya. SeriouslyFish.com (LINK). Electronic version accessed [29/05/2017] (Ref. 115164)</reference>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=118485&amp;speccode=2319'>Offem, B.O., Y.A. Samsons and I.T. Omoniyi0 Length-weight relationship, condition factor and sex ratio of forty six important fishes in a tropical flood river. Res. J. Fish Hydro. 4(2):65-72. (Ref. 118485)</reference>
	<reference isbn='978-3-030-03083-4' url='http://www.fishbase.org/references/FBRefSummary.php?id=120641&amp;speccode=2319'>Skelton, P.H.0 Chapter 11. The freshwater fishes of Angola. p. 207-242. In B.J. Huntley, V. Russo, F. Lages and N. Ferrand (eds.) Biodiversity of Angola. Science &amp; conservation: a modern synthesis. Springer Nature Switzerland AG, Cham, Switzerland. 549 p. (Ref. 120641)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=137577&amp;speccode=2319'>Price, S.A., S.T. Friedman, K.A. Corn, O. Larouche, K. Brockelsby, A.J. Lee, M. Nagaraj, N.G. Bertrand, M. Danao, M.C. Coyne, J.R. Estrada, R. Friedman, E. Hoeft, M. Iwan, D. Gross, J.H. Kao, B. Landry, M.J. Linares, C. McGlinn and …0 FishShapes v1: Functionally relevant measurements of teleost shape and size on three dimensions. Ecology 103(e3829):1-2. (Ref. 137577)</reference><dataObject>
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	<dc:description>An uncommon species which inhabits large rivers (Ref. 4967).  Largest freshwater species in southern Africa; occurs in large deep rivers within the mainstream or in deep pools and lakes or backwaters (Ref. 7248, 78218).  Most active at night, feeding on any available food, including invertebrates and insects when small, fish and other small vertebrates when large; scavenges off large carcasses and offal from riverside villages; live for 12 or more years (Ref. 7248).  Maximum reported size for West Africa and Lower Guinea: 610 mm TL (Ref. 57129, 81643).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=270&amp;speccode=2319'>Teugels, G.G., B. Denayer and M. Legendre 1990 A systematic revision of the African catfish genus &lt;i&gt;Heterobranchus&lt;/i&gt; Geoffroy-Saint-Hilaire, 1809 (Pisces: Clariidae). Zool. J. Linn. Soc. 98:237-257. (Ref. 270)</reference></dataObject><dataObject>
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	<dc:description>Diagnosis: length of rayed dorsal fin 27-36% standard length; 26-35 dorsal-fin rays (Ref. 57129, 81644). Length of adipose fin 24-34% standard length (Ref. 57129, 81644), about equal to the length of the rayed dorsal fin (Ref. 28714). Width of vomerine tooth plate 25.0-32.2%, and width of premaxillary tooth plate 29.5-36.9% head length (Ref. 57129, 81644). Pectoral spine with strong, downward-pointed serrations on the anterior side (Ref. 270, 57129, 81644). Length of pectoral fin 12.7-18.0% SL (mean 16) (Ref. 81644). Posterior part of adipose fin blackish (Ref. 4967, 57129). Caudal fin with a whitish cross-bar (Ref. 57129). Head long, broad and somewhat rectangular in dorsal outline (Ref. 270), its length 28.4-35.9% SL (Ref. 81644). Snout broadly rounded; eyes superolateral in position; frontal fontanelle long and narrow; occipital fontanelle oval-shaped; tooth plates wide; suprabranchial organ well developed; caudal fin barred; openings of secondary canals hardly visible but display a regular pattern (Ref. 270).Description: Coloration: preserved specimens: grey, greyish-brown to dark brown on back and sides, and pale brown to whitish on belly; some individuals marbled, particularly on posterior part of body; caudal fin often crossed by several vertical bars, one of which is whitish, hind margin of fin often white-edged; posterior part of adipose fin darker than anterior part (Ref. 57129, 81644).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=270&amp;speccode=2319'>Teugels, G.G., B. Denayer and M. Legendre 1990 A systematic revision of the African catfish genus &lt;i&gt;Heterobranchus&lt;/i&gt; Geoffroy-Saint-Hilaire, 1809 (Pisces: Clariidae). Zool. J. Linn. Soc. 98:237-257. (Ref. 270)</reference></dataObject><dataObject>
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	<dc:description>Dorsal spines (total): 0; Dorsal soft rays (total): 26 - 35; Analsoft rays: 42 - 52; Vertebrae: 55 - 61</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=270&amp;speccode=2319'>Teugels, G.G., B. Denayer and M. Legendre 1990 A systematic revision of the African catfish genus &lt;i&gt;Heterobranchus&lt;/i&gt; Geoffroy-Saint-Hilaire, 1809 (Pisces: Clariidae). Zool. J. Linn. Soc. 98:237-257. (Ref. 270)</reference></dataObject><dataObject>
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	<dc:description>Africa:  Nile, Congo system, upper and middle Zambezi (Ref. 3820). Reported from Lake Rukwa and the lower Zambezi (Ref. 95585). Also from Lakes Tanganyika and Edward (Ref. 7248). In West Africa known from the Gambia, Upper Senegal, Niger, Benue, Lake Chad, Volta, and coastal basins from Guinea to Nigeria, including the delta of Niger and the Cross river (Ref. 57129). In Lower Guinea present in the Cross and Sanaga in Cameroon and the Ogowe in Gabon (Ref. 81644).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=3820&amp;speccode=2319'>Teugels, G.G. 1986 Clariidae. p. 66-101. In J. Daget, J.-P. Gosse and D.F.E. Thys van den Audenaerde (eds.) Check-list of the freshwater fishes of Africa (CLOFFA). ISNB, Brussels, MRAC, Tervuren; and ORSTOM, Paris. Vol. 2. (Ref. 3820)</reference></dataObject><dataObject>
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	<dc:description>fisheries: minor commercial; aquaculture: commercial; gamefish: yes</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=3126&amp;speccode=2319'>Balon, E.K. 1974 Fishes of Lake Kariba, Africa. T.F.H. Publications, Neptune City, New Jersey. (Ref. 3126)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=7248&amp;speccode=2319'>Skelton, P.H. 1993 A complete guide to the freshwater fishes of southern Africa. Southern Book Publishers. 388 p. (Ref. 7248)</reference>
	<reference isbn='0-7974-0831-4' url='http://www.fishbase.org/references/FBRefSummary.php?id=13337&amp;speccode=2319'>Bell-Cross, G. and J.L. Minshull 1988 The fishes of Zimbabwe. National Museums and Monuments of Zimbabwe, Harare, Zimbabwe. 294 p. (Ref. 13337)</reference></dataObject><dataObject>
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	<dc:description>Potamodromous. Migrating within streams, migratory in rivers, e.g. Saliminus, Moxostoma, Labeo.  Migrations should be cyclical and predictable and cover more than 100 km.</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=51243&amp;speccode=2319'>Riede, K. 2004 Global register of migratory species - from global to regional scales. Final Report of the R&amp;D-Projekt 808 05 081. Federal Agency for Nature Conservation, Bonn, Germany. 329 p. (Ref. 51243)</reference></dataObject><dataObject>
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	<dc:source>longifilis</dc:source>
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	<dc:description>Omnivorous. Inhabits calm waters with muddy and silty substrate (Ref. 40409).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=7248&amp;speccode=2319'>Skelton, P.H. 1993 A complete guide to the freshwater fishes of southern Africa. Southern Book Publishers. 388 p. (Ref. 7248)</reference></dataObject><dataObject>
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	<dc:description>150 cm SL (male/unsexed; (Ref. 4967)); max. published weight: 55.0 kg (Ref. 7248)</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=4967&amp;speccode=2319'>Eccles, D.H. 1992 FAO species identification sheets for fishery purposes. Field guide to the freshwater fishes of Tanzania. Prepared and published with the support of the United Nations Development Programme (project URT/87/016). FAO, Rome. 145 p. (Ref. 4967)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=7248&amp;speccode=2319'>Skelton, P.H. 1993 A complete guide to the freshwater fishes of southern Africa. Southern Book Publishers. 388 p. (Ref. 7248)</reference></dataObject><dataObject>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57073'>IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006.</reference></dataObject><dataObject>
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	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=2319</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Morphology</subject>
	<dc:description>demersal; potamodromous (Ref. 51243); freshwater; depth range 10 - ? m (Ref. 7248)</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=51243&amp;speccode=2319'>Riede, K. 2004 Global register of migratory species - from global to regional scales. Final Report of the R&amp;D-Projekt 808 05 081. Federal Agency for Nature Conservation, Bonn, Germany. 329 p. (Ref. 51243)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=7248&amp;speccode=2319'>Skelton, P.H. 1993 A complete guide to the freshwater fishes of southern Africa. Southern Book Publishers. 388 p. (Ref. 7248)</reference></dataObject>
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