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	<dc:identifier>FB-22566</dc:identifier>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=22566</dc:source>
	<dwc:Kingdom>Animalia</dwc:Kingdom>
	<dwc:Phylum>Chordata</dwc:Phylum>
	<dwc:Class>Elasmobranchii</dwc:Class>
	<dwc:Order>Rajiformes</dwc:Order>
	<dwc:Family>Arhynchobatidae</dwc:Family>
	<dwc:Genus>Arctoraja</dwc:Genus>
	<dwc:ScientificName>Arctoraja simoterus (Ishiyama, 1967)</dwc:ScientificName>
	<commonName xml:lang='en'>Golden skate</commonName>
	<commonName xml:lang='et'>Kuld-süvarai</commonName>
	<commonName xml:lang='ja'>Dobukasube</commonName>
	<commonName xml:lang=''>Goudenrog</commonName>
	<commonName xml:lang=''>斯氏深海鰩</commonName>
	<commonName xml:lang=''>斯氏深海鳐</commonName>
	<synonym relationship='synonym'>Arctoraja smirnovi (Soldatov &amp; Pavlenko, 1915)</synonym>
	<synonym relationship='synonym'>Bathyraja smirnovi smirnovi (Soldatov &amp; Pavlenko, 1915)</synonym>
	<synonym relationship='synonym'>Breviraja smirnovi (Soldatov &amp; Pavlenko, 1915)</synonym>
	<synonym relationship='synonym'>Breviraja smirnovi ankasube Ishiyama, 1958</synonym>
	<synonym relationship='synonym'>Breviraja smirnovi smirnovi (Soldatov &amp; Pavlenko, 1915)</synonym>
	<synonym relationship='synonym'>Raja smirnovi Soldatov &amp; Pavlenko, 1915</synonym>
	<dcterms:created>1996-04-29</dcterms:created>
	<dcterms:modified>2022-01-29</dcterms:modified>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=205&amp;speccode=22567'>Breder, C.M. and D.E. Rosen0 Modes of reproduction in fishes. T.F.H. Publications, Neptune City, New Jersey. 941 p. (Ref. 205)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=559&amp;speccode=22567'>Masuda, H., K. Amaoka, C. Araga, T. Uyeno and T. Yoshino0 The fishes of the Japanese Archipelago. Vol. 1. Tokai University Press, Tokyo, Japan. 437 p. (text). (Ref. 559)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=6876&amp;speccode=22567'>Quast, J.C. and E.L. Hall0 List of fishes of Alaska and adjacent waters with a guide to some of their literature. U.S. Dep. Commer., NOAA Tech. Rep. NMFS SSRF-658, 47 p. (Ref. 6876)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=8630&amp;speccode=22567'>Mould, B.0 A world list of rays. The scientific nomenclature and distribution of the recent Batoidea (Batoidea, Elasmobranchii, Chondrichthyes). University of Nottingham, [UK]. 82 p. (Ref. 8630)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=13360&amp;speccode=22567'>Nakaya, K. and S. Shirai0 Fauna and zoogeography of deep-benthic chondrichthyan fishes around the Japanese Archipelago. Jap. J. Ichthyol. 39(1):37-48. (Ref. 13360)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=27314&amp;speccode=22567'>McEachran, J.D. and K.A. Dunn0 Phylogenetic analysis of skates, a morphologically conservative clade of elasmobranchs (Chondrichthyes: Rajidae). Copeia 1998(2):271-290. (Ref. 27314)</reference>
	<reference isbn='957-8596-59-6' url='http://www.fishbase.org/references/FBRefSummary.php?id=31517&amp;speccode=22567'>Wu, H.L., K.-T. Shao and C.F. Lai (eds.)0 Latin-Chinese dictionary of fishes names. The Sueichan Press, Taiwan. 1028 p. (Ref. 31517)</reference>
	<reference isbn='0-8018-6048-2' url='http://www.fishbase.org/references/FBRefSummary.php?id=35766&amp;speccode=22567'>Compagno, L.J.V.0 Checklist of living elasmobranchs. p. 471-498. In W.C. Hamlett (ed.) Sharks, skates, and rays: the biology of elasmobranch fishes. Johns Hopkins University Press, Maryland. (Ref. 35766)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=38732&amp;speccode=22567'>Anonymous0 Fish collection database of the National Museum of Natural History (Smithsonian Institution). Smithsonian Institution-Division of Fishes. (Ref. 38732)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=41249&amp;speccode=22567'>Ishihara, H.0 Study on the systematics and fishery resources of the North Pacific skates (Pisces: Chondrichthyes: Rajidae). Tokyo University. 186 p. Ph.D. dissertation. (Ref. 41249)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=41250&amp;speccode=22567'>Bor, P.0 Egg-capsules of sharks and skates. www.rajidae.tmfweb.nl/rogtabel.html (Ref. 41250)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=41253&amp;speccode=22567'>Ishiyama, R.0 Fauna Japonica. Rajidae (Pisces). Biogeogr. Soc. Jpn. (Tokyo): i-vi, 1-82. (Ref. 41253)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=43454&amp;speccode=22567'>Bor, P.H.F.0 Nederlandse naamlijst van de recente haaien en roggen (Chondrichthyes: Elasmobranchii) van de wereld. World Wide Web electronic publication www.rajidae.tmfweb.nl, version (05/2002). (Ref. 43454)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=50449&amp;speccode=22567'>Dulvy, N.K. and J.D. Reynolds0 Evolutionary transitions among egg-laying, live-bearing and maternal inputs in sharks and rays. Proc. R. Soc. Lond., Ser. B: Biol. Sci. 264:1309-1315. (Ref. 50449)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=58108&amp;speccode=22567'>Chinese Academy of Fishery Sciences0 Chinese aquatic germplasm resources database. http://zzzy.fishinfo.cn (Ref. 58108)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=83882&amp;speccode=22567'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, March 2010. (Ref. 83882)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=89865&amp;speccode=22567'>Orr, J.W., Stevenson, D.E., Hoff, G.R., Spies, I. and J.D. McEachran0 &lt;i&gt;Bathyraja panthera&lt;/i&gt;, a new species of skate (Rajidae: Arhynchobatinae) from the western Aleutian Islands, and resurrection of the subgenus &lt;i&gt;Arctoraja&lt;/i&gt; Ishiyama. NOAA Professional Paper NMFS 11:50pp (Ref. 89865)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=90062&amp;speccode=22567'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, March 2012. (Ref. 90062)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=95632&amp;speccode=22567'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, April 2014. (Ref. 95632)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=101110&amp;speccode=22567'>FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, [accessed 13/04/2015]. (Ref. 101110)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=105471&amp;speccode=22567'>eElurikkus0 eElurikkus koondab ühte portaali Eesti eluslooduse andmed. http://elurikkus.ut.ee/ (Ref. 105471)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=123274&amp;speccode=22567'>Misawa, R., A.M. Orlov, S.Y. Orlova, I.I. Gordeev, H. Ishihara, T. Hamatsu, Y. Ueda, K. Fujiwara, H. Endo and Y. Kai0 &lt;i&gt;Bathyraja (Arctoraja) sexoculata&lt;/i&gt; sp. nov., a new softnose skate (Rajiformes: Arhynchobatidae) from Simushir Island, Kuril Islands (western North Pacific), with special reference to geographic variations in &lt;i&gt;Bathyraja (Arctoraja) smirnovi&lt;/i&gt;. Zootaxa 4861(4):515-543. (Ref. 123274)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=123477&amp;speccode=22567'>ECoF, Eschmeyer's Catalog of Fishes0 Catalog of Fishes. 1985 - onwards. &lt;b&gt;CITATION NOT TO BE USED OUTSIDE FISHBASE.  SEE REMARKS&lt;/b&gt;. Authorship varies in time: Eschmeyer, W.N.; Fricke, R.; Van der Laan, R.  Web database. http://researcharchive.calacademy.org/research/ichthyology/catalog/fishcatmain.asp (Ref. 123477)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=125494&amp;speccode=22567'>Dyldin, Y.V.0 Annotated checklist of the sharks, batoids and chimaeras (Chondrichthyes: Elasmobranchii, Holocephali) from waters of Russia and adjacent areas. Publ. Seto Mar. Biol. Lab., 43:40-91. (Ref. 125494)</reference>
	<reference isbn='1849954984' url='http://www.fishbase.org/references/FBRefSummary.php?id=128868&amp;speccode=22567'>Beolens, B., M. Grayson and M. Watkins0 Eponym dictionary of Fishes. 	Dunbeath, Caithness (Scotland, UK): Whittles Publishing Limited, 1300 p. (Ref. 128868)</reference>
	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=132125&amp;speccode=22567'>Ferragut-Perello, F., S. Ramirez-Amaro, A.C. Tsikliras, N. Petit-Marty, D. Dimarchopoulou0 Exploitation and conservation status of the thornback ray (&lt;i&gt;Raja clavata&lt;/i&gt;) in the Balearic Islands (Western Mediterranean). Fishes 8(2):117. (Ref. 132125)</reference><dataObject>
	<dc:identifier>FB-GeneralDescr-22567</dc:identifier>
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	<agent homepage='' logoURL='' role='recorder'>Cristina V. Garilao</agent>
	<dcterms:created>1996-04-29 00:00:00</dcterms:created>
	<dcterms:modified>2022-03-02 00:00:00</dcterms:modified>
	<dc:title>Biology</dc:title>
	<dc:language></dc:language>
	<license>http://creativecommons.org/licenses/by-nc/3.0/</license>
	<dc:rights></dc:rights>
	<dcterms:rightsHolder>John E. Randall</dcterms:rightsHolder>
	<dcterms:bibliographicCitation></dcterms:bibliographicCitation>
	<audience>General public</audience>
	<dc:source>http://www.fishbase.org/summary/SpeciesSummary.php?id=22567</dc:source>
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	<dc:description>Oviparous.  Distinct pairing with embrace.  Young may tend to follow large objects, such as their mother (Ref. 205).  Eggs are oblong capsules with stiff pointed horns at the corners deposited in sandy or muddy flats (Ref. 205).  Egg capsules are 12.4-15.0 cm long and 8.7-9.0 cm wide (Ref. 41249, 41253).  Depth range given by H. Ishihara (pers.comm. 07/07).  Reported to feed primarily on amphipods, decapods, fishes, euphausids, and squid on Yamato Bank in the Sea of Japan.  Maximum length reported 116 cm TL (Ref, 89865).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=123274&amp;speccode=22567'>Misawa, R., A.M. Orlov, S.Y. Orlova, I.I. Gordeev, H. Ishihara, T. Hamatsu, Y. Ueda, K. Fujiwara, H. Endo and Y. Kai 2020 &lt;i&gt;Bathyraja (Arctoraja) sexoculata&lt;/i&gt; sp. nov., a new softnose skate (Rajiformes: Arhynchobatidae) from Simushir Island, Kuril Islands (western North Pacific), with special reference to geographic variations in &lt;i&gt;Bathyraja (Arctoraja) smirnovi&lt;/i&gt;. Zootaxa 4861(4):515-543. (Ref. 123274)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Estelita Emily Capuli</agent>
	<dcterms:created>2022-03-02 00:00:00</dcterms:created>
	<dcterms:modified>2022-03-02 00:00:00</dcterms:modified>
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	<dcterms:rightsHolder>John E. Randall</dcterms:rightsHolder>
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	<dc:description>This species is distinguished by the following characters: no blotches on the dorsal disc surface or a pair of white or dark blotches; dark blotch and band usually absent on the ventral disc surface; predorsal caudal vertebrae 80-87 (Ref. 123274).. Geographical variations are present in the populations of the seas of Japan and Okhotsk: number of total tail thorns, ranging from 21-33 (average and standard deviation: 27.3±2.9) (southern Sea of Japan) vs.19-29 (26.0 ± 2.3) (northern Sea of Japan), vs. 22-27 (24.3 ± 1.5) (Sea of Okhotsk) (P &lt; 0.05) and the frequency distribution of tail thorn numbers was bimodal in the Sea of Japan [modes: 25 and 28,29 (southern); 25,26 and 28 (northern)], but unimodal in the Sea of Okhotsk (mode: 25) (Ref. 123274).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=123274&amp;speccode=22567'>Misawa, R., A.M. Orlov, S.Y. Orlova, I.I. Gordeev, H. Ishihara, T. Hamatsu, Y. Ueda, K. Fujiwara, H. Endo and Y. Kai 2020 &lt;i&gt;Bathyraja (Arctoraja) sexoculata&lt;/i&gt; sp. nov., a new softnose skate (Rajiformes: Arhynchobatidae) from Simushir Island, Kuril Islands (western North Pacific), with special reference to geographic variations in &lt;i&gt;Bathyraja (Arctoraja) smirnovi&lt;/i&gt;. Zootaxa 4861(4):515-543. (Ref. 123274)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Cristina V. Garilao</agent>
	<dcterms:created>1996-04-29 00:00:00</dcterms:created>
	<dcterms:modified>2025-11-05 00:00:00</dcterms:modified>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#DiagnosticDescription</subject>
	<dc:description>North Pacific:  Sea of Okhotsk, Kuril Islands, and the Sea of Japan.</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=89865&amp;speccode=22567'>Orr, J.W., Stevenson, D.E., Hoff, G.R., Spies, I. and J.D. McEachran 2011 &lt;i&gt;Bathyraja panthera&lt;/i&gt;, a new species of skate (Rajidae: Arhynchobatinae) from the western Aleutian Islands, and resurrection of the subgenus &lt;i&gt;Arctoraja&lt;/i&gt; Ishiyama. NOAA Professional Paper NMFS 11:50pp (Ref. 89865)</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Susan M. Luna</agent>
	<dcterms:created>1996-04-29 00:00:00</dcterms:created>
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	<audience>General public</audience>
	<dc:source>http://www.fishbase.org//Reproduction/FishReproSummary.php?ID=22567&amp;StockCode=17935</dc:source>
	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#Morphology</subject>
	<dc:description>Oviparous, paired eggs are laid.  Embryos feed solely on yolk (Ref. 50449).  Distinct pairing with embrace. Young may tend to follow large objects, such as their mother (Ref. 205).  Eggs have horn-like projections on the shell (Ref. 205).</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=205&amp;speccode=22567'>Breder, C.M. and D.E. Rosen 1966 Modes of reproduction in fishes. T.F.H. Publications, Neptune City, New Jersey. 941 p. (Ref. 205)</reference></dataObject><dataObject>
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	<dc:title>Max. size</dc:title>
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	<dc:description>102 cm TL (male/unsexed; (Ref. 89865))</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=89865&amp;speccode=22567'>Orr, J.W., Stevenson, D.E., Hoff, G.R., Spies, I. and J.D. McEachran 2011 &lt;i&gt;Bathyraja panthera&lt;/i&gt;, a new species of skate (Rajidae: Arhynchobatinae) from the western Aleutian Islands, and resurrection of the subgenus &lt;i&gt;Arctoraja&lt;/i&gt; Ishiyama. NOAA Professional Paper NMFS 11:50pp (Ref. 89865)</reference></dataObject><dataObject>
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	<dc:description>Least Concern (LC) , IUCN Grouper and Wrasse Specialist Group</dc:description>
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	<reference isbn='' url='http://www.fishbase.org/references/FBRefSummary.php?id=57073'>IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006.</reference></dataObject><dataObject>
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	<agent homepage='' logoURL='' role='recorder'>Cristina V. Garilao</agent>
	<dcterms:created>1996-04-29 00:00:00</dcterms:created>
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	<subject>http://rs.tdwg.org/ontology/voc/SPMInfoItems#TrophicStrategy</subject>
	<dc:description>demersal; marine; depth range 100 - 1000 m</dc:description>
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