FB-16307 http://www.fishbase.org/summary/SpeciesSummary.php?id=16307 Animalia Chordata Teleostei Gobiiformes Gobiidae Millerigobius Millerigobius macrocephalus (Kolombatović, 1891) Nackthalsgrundel Nackthals-Grundel Goad goby Racer goby Rändurmudil Paljaskaulatokko Gobie coureur Bychok-gonets бычок-гонец Küçük kayabalığı Sarıkaya balığı Baboi de glod Guvid Zglavoaca Zlăvoača گاو ماهي 裸喉新鰕虎魚 裸喉新鰕虎鱼 Gobius burmeisteri Kessler, 1877 Gobius gymnotrachelus Kessler, 1857 Gobius macropus De Filippi, 1863 Mesogobius gymnotrachelus (Kessler, 1857) Mesogobius gymnotrachelus otschakovinus Zubovich, 1925 Neogobius gymnotrachelus (Kessler, 1857) 1996-01-19 2014-05-12 Svetovidov, A.N.0 Handbook of the fauna of the USSR, fishes of the Black Sea. Izdatel'stvo Nauka, Moscow, 550 p. (Ref. 897) Berg, L.S.0 Freshwater fishes of the U.S.S.R. and adjacent countries. volume 3, 4th edition. Israel Program for Scientific Translations Ltd, Jerusalem. (Russian version published 1949). (Ref. 2058) Baensch, H.A. and R. Riehl0 Aquarien atlas. Bd. 3. Melle: Mergus, Verlag für Natur-und Heimtierkunde, Germany. 1104 p. (Ref. 2059) Miller, P.J.0 Gobiidae. p. 483-515. In J.C. Hureau and Th. Monod (eds.) Check-list of the fishes of the north-eastern Atlantic and of the Mediterranean (CLOFNAM). UNESCO, Paris. Vol. 1. (Ref. 4345) Hureau, J.-C.0 La base de données GICIM: Gestion informatisée des collections ichthyologiques du Muséum. p. 225-227. In Atlas Préliminaire des Poissons d'Eau Douce de France. Conseil Supérieur de la Pêche, Ministère de l'Environnement, CEMAGREF et Muséum national d'Histoire naturelle, Paris. (Ref. 4517) Miller, P.J.0 Gobiidae. p. 1019-1085. In P.J.P. Whitehead, M.-L. Bauchot, J.-C. Hureau, J. Nielsen and E. Tortonese (eds.) Fishes of the North-eastern Atlantic and the Mediterranean. Volume 3. UNESCO, Paris. (Ref. 4696) Sokolov, L.I. and L.S. Berdicheskii0 Acipenseridae. p. 150-153. In J. Holcík (ed.) The freshwater fishes of Europe. Vol. 1, Part II. General introduction to fishes Acipenseriformes. AULA-Verlag Wiesbaden. 469 p. (Ref. 9953) Baillie, J. and B. Groombridge (eds.)0 1996 IUCN red list of threatened animals. IUCN, Gland, Switzerland. 378 p. (Ref. 12255) Vasil'ev, V.P. and K.A. Grogoryan0 Karyology of fishes from the family Gobiidae. Vopr. Ikhtiol. 32(5):27-40. (Ref. 12380) Kottelat, M.0 European freshwater fishes. An heuristic checklist of the freshwater fishes of Europe (exclusive of former USSR), with an introduction for non-systematists and comments on nomenclature and conservation. Biologia, Bratislava, 52/Suppl. 5:1-271. (Ref. 13696) Reshetnikov, Y.S., N.G. Bogutskaya, E.D. Vasil'eva, E.A. Dorofeeva, A.M. Naseka, O.A. Popova, K.A. Savvaitova, V.G. Sideleva and L.I. Sokolov0 An annotated check-list of the freshwater fishes of Russia. J. Ichthyol. 37(9):687-736. (Ref. 26334) Wu, H.L., K.-T. Shao and C.F. Lai (eds.)0 Latin-Chinese dictionary of fishes names. The Sueichan Press, Taiwan. 1028 p. (Ref. 31517) Anonymous0 Fish collection database of the Natural History Museum, London (formerly British Museum of Natural History (BMNH)). Natural History Museum, London (formerly British Museum of Natural History (BMNH)). (Ref. 31982) Klinkhardt, M, M. Tesche and H. Greven0 Database of fish chromosomes. Westarp Wissenschaften. (Ref. 34370) Hilton-Taylor, C.0 2000 IUCN red list of threatened species. IUCN, Gland, Switzerland and Cambridge, UK. xviii + 61 p. (with 1 CD-ROM). (Ref. 36508) Anonymous0 Fish collection database of the National Museum of Natural History (Smithsonian Institution). Smithsonian Institution-Division of Fishes. (Ref. 38732) Anonymous0 Fish collection database of the Zoological Museum, University of Copenhagen. Zoological Museum, University of Copenhagen. (Ref. 40919) Anonymous0 Fish collection database of the American Museum of Natural History. American Museum of Natural History, Central Park West , NY 10024-5192, USA. (Ref. 41414) Deval, M.C.0 Common names of marine fishes of Turkey. Unpublished report. (Ref. 41476) Bogutskaya, N.G. and A.M. Naseka0 An overview of nonindigenous fishes in inland waters of Russia. Proc. Zool. Inst. Russ. Acad. Sci. 296:21-30. (Ref. 45022) Bogutskaya, N.G.0 Common names of Russian freshwater fishes. Spreadsheet with cyrillic names made available for inclusion in FishBase. Unpublished. (Ref. 45043) Salekhova, L.P., N.S. Kostenko, T.A. Bogachick and O.N. Minibaeva0 Composition of ichthyofauna in the region of the Karadag State Reserve (Black Sea). J. Ichthyol. 28(2):16-23. (Ref. 45714) Anonymous0 Fish collection of the Royal Ontario Museum. Royal Ontario Museum. (Ref. 47438) Reshetnikov, Y.S.0 Atlas of Russian freshwater fishes, Vol. 2. Moscow Nauka. (Ref. 50519) Louisy, P.0 Guide d'identification des poissons marins. Europe et Méditerranée. Paris: Eds. Eugène Ulmer. (Ref. 50675) Hureau, J.-C. and T. Monod (eds.)0 Supplement. Check-list of the fishes of the north-eastern Atlantic and of the Mediterranean. p.339-394. In J.-C. Hureau and Th. Monod (eds.) Check-list of the fishes of the north-eastern Atlantic and of the Mediterranean.United Nations Educational Scientific and Cultural Organization (UNESCO), Paris, France. Vols 1-2. 683 p. (Ref. 51024) Varjo, M., L. Koli and H. Dahlström0 Kalannimiluettelo (versio 10/03). Suomen Biologian Seura Vanamo Ry. (Ref. 51086) Wiesner, C.0 New records of non-indigenous gobies (<i>Neogobius</i> sp.) in the Austrian Danube. p. 71-72 in Abstracts of the XI European Congress of Ichthyology (ECI XI). Estonian Marine Institute Report Series No. 12. (Ref. 52471) Grabowska, J. and M. Grabowska0 "How the west was won" - expansion of the Ponto-Caspian gobiid fishes. p. 76 in Abstracts of the XI European Congress of Ichthyology (ECI XI). Estonian Marine Institute Report Series No. 12. (Ref. 52518) Solarz, W.0 Alien Species in Poland. Institute of Nature Conservation, Polish Academy of Sciences website. (Ref. 55773) Simonovic, P.0 Ribe Srbije. Zavod za zastitu prirode Srbije i Bioloski faultet Univerziteta u Beogradu. Belgrade. 1247 p. (Ref. 56371) Jankovic, D. and J. Cetkovic-Krpo0 Diverzitet slatkovodnih riba (Osteichthyes) i kolousta (Cephalaspidomorpha) Jugoslavije sa pregledom vrsta od medjunarodnog znacaja. p. 425-445. In V. Stevanovic and V. Vasic (eds) Biodiverzitet Jugoslavije. Bioloski faultet i Ekolibri. Belgrade. (Ref. 56374) Simonovic, P.D.0 Phylogenetic relationships of Ponto-Caspian gobies and their relationship to the Atlantic-Mediterranean Gobiinae. J. Fish Biol. 54(3):533-555. (Ref. 56375) Hegedis, A., M. Nikcevic, B. Mickovic, D. Jankovic and R.K. Andjus0 Discovery of the goby <i>Neogobius gymnotrachelus</i> in Yugoslav fresh waters. Arhiv Bioloskih Nauka 43(3-4):39-40. (Ref. 56376) Iacob, L. and I. Dima0 Regnul Animalia. Eukarya. Enciclopedia Florei si Faunei din Romania. (Ref. 56616) Chinese Academy of Fishery Sciences0 Chinese aquatic germplasm resources database. http://zzzy.fishinfo.cn (Ref. 58108) Bogutskaya, N.G. and A.M. Naseka0 REGIONAL CHECK-LISTS: Don River Drainage Area. In Website and Database: "Freshwater Fishes of Russia": A Source of Information on the Current State of the Fauna. Zoological Institute RAS. (Ref. 58293) Bogutskaya, N.G. and A.M. Naseka0 REGIONAL CHECK-LISTS: Kuban River Drainage Area. In Website and Database: "Freshwater Fishes of Russia": A Source of Information on the Current State of the Fauna. Zoological Institute RAS. (Ref. 58296) Bogutskaya, N.G. and A.M. Naseka0 REGIONAL CHECK-LISTS: Volga River Drainage Area. In Website and Database: "Freshwater Fishes of Russia": A Source of Information on the Current State of the Fauna. Zoological Institute RAS. (Ref. 58297) Fricke, R., M. Bilecenoglu and H.M. Sari0 Annotated checklist of fish and lamprey species (Gnathostoma and Petromyzontomorphi) of Turkey, including a Red List of threatened and declining species. Stuttgarter Beitr. Naturk. Sea A (706):1-172. (Ref. 58342) Grabowska, J.0 Reproductive biology of racer goby (<i>Neogobius gymnotrachelus</i> in the Wloclawski Reservoir (Vistula River, Poland). J. Appl. Ichthyol. 21:296-299. (Ref. 58957) Kottelat, M. and J. Freyhof0 Handbook of European freshwater fishes. Publications Kottelat, Cornol and Freyhof, Berlin. 646 pp. (Ref. 59043) Tarkan, A.S., O. Gaygusuz, H. Acipinar, C. Gürsoy and M. Ozulug0 Length-weight relationship of fishes from the Marmara region (NW-Turkey). J. Appl. Ichthyol. 22:271-273. (Ref. 59105) Tsykhon-Lukanina, E.A.0 Feeding of gobies of the northern Caspian Sea. Trudy Vsesoyuznogo Gidrobiologicheskogo Obshchestva 9:214-239. (Ref. 75130) Neilson, M.E. and C.A. Stepien0 Escape from the Ponto-Caspian: Evolution and biogeography of an endemic goby species flock (Benthophilinae: Gobiidae: Teleostei). Molecular Phylogenetics and Evolution, 52(2009):84-102. (Ref. 83713) AFMAFEWM (Austrian Federal Ministry of Agriculture, Forestry, Environment and Water Management)0 Checklist of species in Austria. Federal Office for Water Management, Institute of Waters Ecology, Fisheries and Lake Research. (Ref. 87703) Borcherding, J., S. Gertzen and S. Staas0 First record of Pontian racer goby, <i>Babka gymnotrachelus</i> (Gobiidae: Teleostei), in the River Rhine, Germany. J. Appl. Ichthyol. 27(6):1399-1400. (Ref. 88692) Patzner, R.A., J.L. Van Tassell, M. Kovačić and B.G. Kapoor0 The biology of gobies. Enfield, NH : Science Publishers ; Boca Raton, FL : DIstributed by CRC Press, 685 p. (Ref. 92840) Grabowska, J., J. Kotusz and A. Witkowski0 Alien invasive fish species in Polish waters: an overview. Folia Zool. 59(1):73-85. (Ref. 92885) eElurikkus0 eElurikkus koondab ühte portaali Eesti eluslooduse andmed. http://elurikkus.ut.ee/ (Ref. 105471) Çiçek, E., R. Fricke and S.S. Birecikligil0 Freshwater fishes of Turkey: a revised and updated annotated checklist. Biharean Biologist 9(2):141-157. (Ref. 106151) Semenchenko, V.P., V.K. Rizevsky, S.E. Mastitsky, V.V. Vezhnovets, M.V. Pluta, V.I. Razlutsky and T. Laenko0 Checklist of aquatic alien species established in large river basins of Belarus. Aquatic Invasions 4(2): 337-347. (Ref. 115062) IUCN0 The IUCN Red List of Threatened Species. 1964 - onwards. https://www.iucnredlist.org/ (Ref. 130435) FB-GeneralDescr-16309 http://purl.org/dc/dcmitype/Text text/html Pascualita Sa-a 1996-01-19 00:00:00 2024-07-18 00:00:00 Biology http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=16309 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Size Occurs in brackish- and fresh-water habitats with low salinity (< 2 ppt); lagoons and lakes; large rivers to small, fast-flowing streams; on sand or mud bottom; mainly in well vegetated or high-complexity habitats. Abundant in backwaters and still channels (Ref. 59043). Longevity is 4-5 years; spawns for the first time at 2 years; spawning season in April to June, occasionally until mid-August; females may repeat spawning during a season; usually spawns for a single season. Males guard eggs until hatching; with adhesive eggs deposited on stones, shells and aquatic plants (Ref. 59043). Feeds on crustaceans (esp. Corophiid amphipods), aquatic insects (mostly chironomid larvae), polychaetes, also small fish and mollusks (Ref. 4696). Miller, P.J. 1986 Gobiidae. p. 1019-1085. In P.J.P. Whitehead, M.-L. Bauchot, J.-C. Hureau, J. Nielsen and E. Tortonese (eds.) Fishes of the North-eastern Atlantic and the Mediterranean. Volume 3. UNESCO, Paris. (Ref. 4696) FB-DiagnosticDescr-16309 http://purl.org/dc/dcmitype/Text text/html Rainer Froese 2004-02-10 00:00:00 2013-01-15 00:00:00 http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=16309 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Threats This species is distinguished from its congeners entering freshwater in Europe by the following characters: irregular position and shape of diagonal bars on body; first branched ray of second dorsal about as long as penultimate ray; no scales on midline of nape, in front of preoperculum; pelvic-disc fraenum with small rounded lobes and the length is less than 1/6 of width at base; scales in midlateral series 54-62 + 2-3; posterior part of first dorsal without black spot (Ref. 59043). Reshetnikov, Y.S. 2003 Atlas of Russian freshwater fishes, Vol. 2. Moscow Nauka. (Ref. 50519) FB-Morphology-16309 http://purl.org/dc/dcmitype/Text text/html Rainer Froese 2004-02-10 00:00:00 2013-01-15 00:00:00 http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=16309 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Habitat Dorsal spines (total): 7 - 8; Dorsal soft rays (total): 14 - 18; Analspines: 1; Analsoft rays: 12 - 16 Reshetnikov, Y.S. 2003 Atlas of Russian freshwater fishes, Vol. 2. Moscow Nauka. (Ref. 50519) FB-Distribution-16309 http://purl.org/dc/dcmitype/Text text/html Pascualita Sa-a 1996-01-19 00:00:00 2025-11-05 00:00:00 http://creativecommons.org/licenses/by-nc/3.0/ John E. Randall General public http://www.fishbase.org/summary/SpeciesSummary.php?id=16309 Eurasia: Widespread all over the Marmara, Black and Caspian seas basins; inhabits mostly fresh- and brackish waters with low salinity ( Patzner, R.A., J.L. Van Tassell, M. Kovačić and B.G. Kapoor 2011 The biology of gobies. Enfield, NH : Science Publishers ; Boca Raton, FL : DIstributed by CRC Press, 685 p. (Ref. 92840) FB-Uses-16309 http://purl.org/dc/dcmitype/Text text/html Pascualita Sa-a 1996-01-19 00:00:00 2024-07-18 00:00:00 Importance http://creativecommons.org/licenses/by-nc/3.0/ John E. Randall General public http://www.fishbase.org/summary/SpeciesSummary.php?id=16309 fisheries: of no interest FB-LifeCycle-16309 http://purl.org/dc/dcmitype/Text text/html Armi G. Torres 1996-01-19 00:00:00 2024-07-18 00:00:00 http://creativecommons.org/licenses/by-nc/3.0/ John E. Randall General public http://www.fishbase.org//Reproduction/FishReproSummary.php?ID=16309&StockCode=15037 Adhesive eggs are deposited on stones, shells and aquatic plants. Males guard eggs until hatching (Ref.59043). Sokolov, L.I. and L.S. Berdicheskii 1989 Acipenseridae. p. 150-153. In J. Holcík (ed.) The freshwater fishes of Europe. Vol. 1, Part II. General introduction to fishes Acipenseriformes. AULA-Verlag Wiesbaden. 469 p. (Ref. 9953) FB-TrophicStrategy-16309 http://purl.org/dc/dcmitype/Text text/html Drina Sta. Iglesia 2001-01-17 00:00:00 2008-04-15 00:00:00 http://creativecommons.org/licenses/by-nc/3.0/ John E. Randall General public gymnotrachelus Occurs in lower reaches of rivers and coastal lagoons, rarely mesohaline; on sand or mud. Feeds on crustaceans (esp. corophiid amphipods), aquatic insects (mostly chironomid larvae), polychaetes, also small fish and molluscs. Miller, P.J. 1986 Gobiidae. p. 1019-1085. In P.J.P. Whitehead, M.-L. Bauchot, J.-C. Hureau, J. Nielsen and E. Tortonese (eds.) Fishes of the North-eastern Atlantic and the Mediterranean. Volume 3. UNESCO, Paris. (Ref. 4696) FB-Size-16309 http://purl.org/dc/dcmitype/Text text/html Pascualita Sa-a 1996-01-19 00:00:00 2024-07-18 00:00:00 Max. size http://creativecommons.org/licenses/by-nc/3.0/ John E. Randall General public http://www.fishbase.org/summary/SpeciesSummary.php?id=16309 16.2 cm TL (male/unsexed; (Ref. 2058)); max. reported age: 5 years (Ref. 59043) Berg, L.S. 1965 Freshwater fishes of the U.S.S.R. and adjacent countries. volume 3, 4th edition. Israel Program for Scientific Translations Ltd, Jerusalem. (Russian version published 1949). (Ref. 2058) Kottelat, M. and J. Freyhof 2007 Handbook of European freshwater fishes. Publications Kottelat, Cornol and Freyhof, Berlin. 646 pp. (Ref. 59043) FB-Threats-2 http://purl.org/dc/dcmitype/Text text/html Belen Acosta 1990-10-18 00:00:00 2024-03-07 00:00:00 http://creativecommons.org/licenses/by-nc/3.0/ John E. Randall General public http://www.fishbase.org/summary/SpeciesSummary.php?id=2 Least Concern (LC) , IUCN Grouper and Wrasse Specialist Group IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006. FB-Habitat-16309 http://purl.org/dc/dcmitype/Text text/html Pascualita Sa-a 1996-01-19 00:00:00 2024-07-18 00:00:00 Environment http://creativecommons.org/licenses/by-nc/3.0/ John E. Randall General public http://www.fishbase.org/summary/SpeciesSummary.php?id=16309 benthopelagic; freshwater; brackish