FB-10741 http://www.fishbase.org/summary/SpeciesSummary.php?id=10741 Animalia Chordata Teleostei Characiformes Lebiasinidae Copella Copella arnoldi (Regan, 1912) Smuklen pryskacz Pirá tantan Piratanta Sprøjtekarpelaks Spritzsalmler Copeina Jumping characin Splash tetra Splashing samlet Splashing tetra Spotted characin Spraying characin Arnoldi kopella Roiskuttajatetra Viirisukkulatetra Stänktetra Milé montangn Milipipi Mulet montagne Pilamagnoe-magnone Salmovka brizgačka Ti-yaya 圭亚那丝鳍脂鲤 圭亞那絲鰭脂鯉 阿氏短頜鮰脂鯉 阿氏短颌鮰脂鲤 阿氏絲鰭脂鯉 阿氏丝鳍脂鲤 Copeina arnoldi Regan, 1912 Copeina carsevennensis Regan, 1912 Copeina eigenmanni (non Regan, 1912) Copella carsevennensis (Regan, 1912) Copella eigenmanni (non Regan, 1912) Copella metae (non Eigenmann, 1914) Copella nattereri (non Steindachner, 1876) 1994-06-14 2022-05-16 Breder, C.M. and D.E. Rosen0 Modes of reproduction in fishes. T.F.H. Publications, Neptune City, New Jersey. 941 p. (Ref. 205) Géry, J.0 Characoids of the world. Neptune City ; Reigate : T.F.H. [etc.]; 672 p. : ill. (chiefly col.) ; 23 cm. (Ref. 598) Riehl, R. and H.A. Baensch0 Aquarien Atlas. Band. 1. Melle: Mergus, Verlag für Natur-und Heimtierkunde, Germany. 992 p. (Ref. 1672) Robins, C.R., R.M. Bailey, C.E. Bond, J.R. Brooker, E.A. Lachner, R.N. Lea and W.B. Scott0 World fishes important to North Americans. Exclusive of species from the continental waters of the United States and Canada. Am. Fish. Soc. Spec. Publ. (21):243 p. (Ref. 4537) Grabda, E. and T. Heese0 Polskie nazewnictwo popularne kraglouste i ryby. Cyclostomata et Pisces. Wyzsza Szkola Inzynierska w Koszalinie. Koszalin, Poland. 171 p. (in Polish). (Ref. 6313) Mills, D. and G. Vevers0 The Tetra encyclopedia of freshwater tropical aquarium fishes. Tetra Press, New Jersey. 208 p. (Ref. 7020) Porto, J.I.R., E. Feldberg, C.M. Nakayama and J.N. Falcao0 A checklist of chromosome numbers and karyotypes of Amazonian freshwater fishes. Rev. Hydrobiol. Trop. 25(4):287-299. (Ref. 7052) Pereira, R.0 Peixes de nossa terra. Livraria Nobel, Sao Paulo, Brazil. 129 p. (Ref. 7145) Balon, E.K.0 Epigenesis of an epigeneticist: the development of some alternative concepts on the early ontogeny and evolution of fishes. Guelph Ichthyol. Rev. 1:1-48. (Ref. 7471) Kenny, J.S.0 Views from the bridge: a memoir on the freshwater fishes of Trinidad. Julian S. Kenny, Maracas, St. Joseph, Trinidad and Tobago. 98 p. (Ref. 11225) Planquette, P., P. Keith and P.-Y. Le Bail0 Atlas des poissons d'eau douce de Guyane. Tome 1. Collection du Patrimoine Naturel Volume 22, MNHN, Paris & INRA, Paris. 429 p. (Ref. 12225) Eschmeyer, W.N.0 PISCES. Eschmeyer's PISCES database as published on the Internet in June 1997, URL: gopher://gopher.calacademy.org:640/7. (Ref. 12965) Riehl, R. and H.A. Baensch0 Aquarien Atlas, Band 1. 10th edition. Mergus Verlag GmBH, Melle, Germany. 992 p. (Ref. 13371) Boujard, T., M. Pascal, F.J. Meunier and P.-Y. Le Bail0 Poissons de Guyane. Guide écologique de l'Approuague et de la réserve des Nouragues. Institut National de la Recherche Agronomique, Paris, 219 p. (Ref. 27188) Schliewen, U.K.0 Aquarium fish. Barron's Education Series, Incorporated. 159 p. (Ref. 30491) Wu, H.L., K.-T. Shao and C.F. Lai (eds.)0 Latin-Chinese dictionary of fishes names. The Sueichan Press, Taiwan. 1028 p. (Ref. 31517) Anonymous0 Fish collection database of the Natural History Museum, London (formerly British Museum of Natural History (BMNH)). Natural History Museum, London (formerly British Museum of Natural History (BMNH)). (Ref. 31982) Mérigoux, S. and D. Ponton0 Body shape, diet and ontogenetic diet shifts in young fish of the Sinnamary River, French Guiana, South America. J. Fish Biol. 52(3):556-569. (Ref. 32839) Klinkhardt, M, M. Tesche and H. Greven0 Database of fish chromosomes. Westarp Wissenschaften. (Ref. 34370) Scheel, J.J.0 Fish chromosomes and their evolution. Interval Report of Danmarks Akvarium, Charlottenlund, Denmark. (Ref. 35080) Anonymous0 The icthyological collection of the Zoological Museum Hamburg (ZMH). Division of Ichthyology and Herpetology, Zoological Museum Hamburg (ZMH). (Ref. 35508) Weitzman, M. and S.H. Weitzman0 Lebiasinidae (Pencil fishes). p. 241-251. In R.E. Reis, S.O. Kullander and C.J. Ferraris, Jr. (eds.) Checklist of the Freshwater Fishes of South and Central America. Porto Alegre: EDIPUCRS, Brasil. (Ref. 37104) Anonymous0 Fish collection database of the National Museum of Natural History (Smithsonian Institution). Smithsonian Institution-Division of Fishes. (Ref. 38732) Anonymous0 Fish collection database of the Zoological Museum, University of Copenhagen. Zoological Museum, University of Copenhagen. (Ref. 40919) Anonymous0 Fish collection database of the American Museum of Natural History. American Museum of Natural History, Central Park West , NY 10024-5192, USA. (Ref. 41414) Anonymous0 Fish collection of the Royal Ontario Museum. Royal Ontario Museum. (Ref. 47438) Varjo, M., L. Koli and H. Dahlström0 Kalannimiluettelo (versio 10/03). Suomen Biologian Seura Vanamo Ry. (Ref. 51086) Carl, H.0 Danish fish names. Zoological Museum of Copenhagen. Unpublished. (Ref. 51471) Ponton, D. and S. Mérigoux0 Description and ecology of some early life stages of fishes in the River Sinnamary (French Guiana, South America). Folia Zool. 50(Monogr. 1):1-116. (Ref. 52185) Chinese Academy of Fishery Sciences0 Chinese aquatic germplasm resources database. http://zzzy.fishinfo.cn (Ref. 58108) FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis, 29 April 2008. (Ref. 75549) FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, March 2010. (Ref. 83882) Nomura, H.0 Nomes científicos dos peixes e seus correspondentes nomes vulgares. In H. Nomura (ed.). Dicionário dos peixes do Brasil. Editerra, Brasília, Brasil: 27-63. (Ref. 86306) FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, March 2012. (Ref. 90062) Brosse, S., G. Grenouillet, M. Gevrey, K. Khazraie and L. Tudesque0 Small-scale gold mining erodes fish assemblage structure in small neotropical streams. Biodivers Conserv (2011)20:1013-1026. (Ref. 90865) FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, April 2014. (Ref. 95632) FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en, [accessed 13/04/2015]. (Ref. 101110) Pihu, E.0 Loomade elu 4. köide. Kalad. Pihu, E. 1979. Loomade Elu, 4. Köide, Kalad. Valgus, Tallinn. (Ref. 105472) FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from http://www.fao.org/fishery/collection/asfis/en (accessed 08/06/2017). (Ref. 115257) Nobile, A.B., D. Freitas-Souza, F.P. Lima, L.B. Vieira, B.F. Melo and C. Oliveira0 Length-weight and length-length relationships of 16 fish species from Amapá, Brazilian Amazon. J. Appl. Ichthyol. 33:1058-1061. (Ref. 116983) FAO-FIES0 Aquatic Sciences and Fisheries Information System (ASFIS) species list. Retrieved from htttps://www.fao.org/fishery/collection/asfis/en [accessed May, 2018] (Ref. 118272) Freitas, T.M.S., A.T.T. De Paula, H. Leão, N.L. Benone and L.F.A. Montag0 Length-weight relationships of 11 fish species from streams of Anapu River Basin, State of Pará, eastern Amazon, Brazil. J. Appl. Ichthyol. 35:793-795. (Ref. 124474) Machado, A.F.d.V.N., C.M.C. Lobato, R.R. Gusmão, L.F.d.A. Montag and B.S. Prudente0 Length-weight relationships of eleven fish species captured in 18 streams of the Capim River basin, Brazil. J. Appl. Ichthyol. 36:745-747. (Ref. 125972) Marčeta, B. and K. Pliberšek (eds)0 BiosWeb. Biological database of the Fisheries Research Institute of Slovenia. Ljubljana, Fisheries Research Institute of Slovenia, 2014. (www.biosweb.org. updated 13/04/2023). (Ref. 128520) Beolens, B., M. Grayson and M. Watkins0 Eponym dictionary of Fishes. Dunbeath, Caithness (Scotland, UK): Whittles Publishing Limited, 1300 p. (Ref. 128868) Marinho, M.M.F. and N.A. Menezes0 Taxonomic review of <i>Copella</i> (Characiformes: Lebiasinidae) with an identification key for the species. PLoS ONE 12(8):e0183069:1-53. (Ref. 137764) FB-GeneralDescr-10741 http://purl.org/dc/dcmitype/Text text/html Armi G. Torres 1994-06-14 00:00:00 2025-08-14 00:00:00 Biology http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=10741 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Morphology Occurs in groups at the surface of small creeks with clear waters. Also found in ponds where it is found together with Rivulus (Ref. 27188). Feeds on Ephemeroptera larvae and ants (Ref. 12225); also on worms, insects and crustaceans (Ref. 7020). Known among aquarists by its unique breeding behavior and parental care: male and the female line up side by side at the surface of the water and jump together out of the water, to spawn. The fertilized eggs are then laid on the underside of an emergent leaf and the male then splashes them with its tail for about three days until they hatch, hence the popular name 'Splash tetra'. It was also observed that just before lining up to jump out of the water toward the upper glass of the aquarium, the female follows the male, touching her abdomen at the anterodorsal portion of the male severaltimes, swimming agitatedly (Ref. 137764). The spawned eggs are deposited on submerged leaves and are guarded by the males (Ref. 12225). During incubation, the male oxygenates the eggs (Ref. 27188). Marinho, M.M.F. and N.A. Menezes 2017 Taxonomic review of <i>Copella</i> (Characiformes: Lebiasinidae) with an identification key for the species. PLoS ONE 12(8):e0183069:1-53. (Ref. 137764) FB-DiagnosticDescr-10741 http://purl.org/dc/dcmitype/Text text/html Estelita Emily Capuli 2025-08-14 00:00:00 2025-08-14 00:00:00 http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=10741 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution This species is distinguished from all its congeners, except some specimens of C. nattereri by the procurrent caudal-fin rays that are hyaline (vs. black); further differs from C. nattereri by the absence of a black mark on each body scale (vs. presence); further differs from all its congeners by having a pigmented area extending anterodorsally from ventral tip of the dentary to ventral portion of the eye (vs. absent). Colouration: some males are unique in having brilliant white spots on scales of the third, fourth, fifth, and sixth longitudinal scale rows (Ref. 137764). Marinho, M.M.F. and N.A. Menezes 2017 Taxonomic review of <i>Copella</i> (Characiformes: Lebiasinidae) with an identification key for the species. PLoS ONE 12(8):e0183069:1-53. (Ref. 137764) FB-Morphology-10741 http://purl.org/dc/dcmitype/Text text/html Estelita Emily Capuli 2025-08-14 00:00:00 2025-08-14 00:00:00 http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=10741 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Uses Dorsal soft rays (total): 8; Analsoft rays: 9; Vertebrae: 37 Marinho, M.M.F. and N.A. Menezes 2017 Taxonomic review of <i>Copella</i> (Characiformes: Lebiasinidae) with an identification key for the species. PLoS ONE 12(8):e0183069:1-53. (Ref. 137764) FB-Distribution-10741 http://purl.org/dc/dcmitype/Text text/html Armi G. Torres 1994-06-14 00:00:00 2025-08-14 00:00:00 http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=10741 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Size South America: lower rio Amazonas basin, coastal drainages of Pará and Amapá, Brazil,Guyana, French Guiana, Surinam, mouth of rio Orinoco, and coastal drainages of Sucre and Monagas, Venezuela. Marinho, M.M.F. and N.A. Menezes 2017 Taxonomic review of <i>Copella</i> (Characiformes: Lebiasinidae) with an identification key for the species. PLoS ONE 12(8):e0183069:1-53. (Ref. 137764) FB-Uses-10741 http://purl.org/dc/dcmitype/Text text/html Armi G. Torres 1994-06-14 00:00:00 2025-08-14 00:00:00 Importance http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=10741 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Threats fisheries: commercial; aquarium: commercial Mills, D. and G. Vevers 1989 The Tetra encyclopedia of freshwater tropical aquarium fishes. Tetra Press, New Jersey. 208 p. (Ref. 7020) Mills, D. and G. Vevers 1989 The Tetra encyclopedia of freshwater tropical aquarium fishes. Tetra Press, New Jersey. 208 p. (Ref. 7020) FB-LifeCycle-10741 http://purl.org/dc/dcmitype/Text text/html Armi G. Torres 1994-06-14 00:00:00 2025-08-14 00:00:00 http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public http://www.fishbase.org//Reproduction/FishReproSummary.php?ID=10741&StockCode=11064 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Habitat Deposits eggs on underside of objects above the water surface. Male in water splashes the clutch periodically (Ref. 7471). In the aquarium, male and female swim vertically to the surface, flick their tails and leap up to a leaf. Female lays from 5 to 8 eggs on the leaf and male fertilizes them immediately. This procedure is repeated several times until some hundreds of eggs have been laid. Male then keeps the eggs damp by flicking his tail to spray them with water. As eggs hatch, fry fall into the water (Ref. 7020). Balon, E.K. 1990 Epigenesis of an epigeneticist: the development of some alternative concepts on the early ontogeny and evolution of fishes. Guelph Ichthyol. Rev. 1:1-48. (Ref. 7471) FB-TrophicStrategy-10741 http://purl.org/dc/dcmitype/Text text/html Drina Sta. Iglesia 2000-12-19 00:00:00 2025-08-14 00:00:00 http://creativecommons.org/licenses/by-nc/3.0/ FAO General public arnoldi Occurs in groups at the surface of small creeks with clear waters. Also found in ponds where it is found together with Rivulus (Ref. 27188). Feeds on Ephemeroptera larvae and ants (Ref. 12225); also on worms, insects and crustaceans (Ref. 7020). Known among aquarists by its unique breeding behavior and parental care: male and the female line up side by side at the surface of the water and jump together out of the water, to spawn. The fertilized eggs are then laid on the underside of an emergent leaf and the male then splashes them with its tail for about three days until they hatch, hence the popular name 'Splash tetra'. It was also observed that just before lining up to jump out of the water toward the upper glass of the aquarium, the female follows the male, touching her abdomen at the anterodorsal portion of the male several times, swimming agitatedly (Ref. 137764). Mills, D. and G. Vevers 1989 The Tetra encyclopedia of freshwater tropical aquarium fishes. Tetra Press, New Jersey. 208 p. (Ref. 7020) FB-Size-10741 http://purl.org/dc/dcmitype/Text text/html Armi G. Torres 1994-06-14 00:00:00 2025-08-14 00:00:00 Max. size http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=10741 http://rs.tdwg.org/ontology/voc/SPMInfoItems#DiagnosticDescription 4.2 cm SL (male/unsexed; (Ref. 137764)); 3.25 cm SL (female); max. published weight: 0.3 g (Ref. 125972) Marinho, M.M.F. and N.A. Menezes 2017 Taxonomic review of <i>Copella</i> (Characiformes: Lebiasinidae) with an identification key for the species. PLoS ONE 12(8):e0183069:1-53. (Ref. 137764) Machado, A.F.d.V.N., C.M.C. Lobato, R.R. Gusmão, L.F.d.A. Montag and B.S. Prudente 2020 Length-weight relationships of eleven fish species captured in 18 streams of the Capim River basin, Brazil. J. Appl. Ichthyol. 36:745-747. (Ref. 125972) FB-Threats-2 http://purl.org/dc/dcmitype/Text text/html Belen Acosta 1990-10-18 00:00:00 2024-03-07 00:00:00 http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=2 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Morphology Least Concern (LC) , IUCN Grouper and Wrasse Specialist Group IUCN 2006 2006 IUCN red list of threatened species. www.iucnredlist.org. Downloaded July 2006. FB-Habitat-10741 http://purl.org/dc/dcmitype/Text text/html Armi G. Torres 1994-06-14 00:00:00 2025-08-14 00:00:00 Environment http://creativecommons.org/licenses/by-nc/3.0/ WorldFish Center - FishBase General public http://www.fishbase.org/summary/SpeciesSummary.php?id=10741 http://rs.tdwg.org/ontology/voc/SPMInfoItems#Distribution benthopelagic; freshwater; pH range: 6.0 - 8.0; dH range: 5 - 12