Issue   
The synonym P. quadrispinosa (Weber, 1913) is valid and redescriberd according to Johnson & White, 2012 (Ref. 90268).
			
			
				
					Environment: milieu / climate zone / depth range / distribution range					
						Ecology					
				
				
				
					Marine;  reef-associated; depth range 0 - 100 m (Ref. 128797). Tropical; 30°N -   24°S				
				 
			
			
			
				
				
				
					Indo-West Pacific:  India and Sri Lanka to Tonga (Ref. 53797); , Ryukyu Islands and the Great Barrier Reef to the Marshall Islands and Samoa.
				
				 
			
			
				
					Size / Weight / Age
				
				
				
					Maturity: Lm ?  range ? - ? cm
 Max length : 24.0 cm TL male/unsexed; (Ref. 48636)				
				 
			
			
						
				
				
				
					Dorsal spines (total): 4 - 5; Dorsal soft rays (total): 20 - 21; Anal spines: 1; Anal soft rays: 17.  A row of small blackish spots in outer third of anal fin (Ref 42740).  Males develop a conspicuous ocellus on each side of the nape (Ref. 1602).  Young are best identified by the patch of small spots on the cheek that usually remains evident in adults (Ref. 48636).
Body shape (shape guide): elongated; Cross section: oval.				
				 
			
			
			
			
			
			
			
				Inhabits both clear lagoon and seaward reefs, in areas of open sand and rubble as well as on rocky surfaces between coral heads.  Also in current prone channels (Ref. 48636).  May be found solitary or form small groups (Ref. 90102).			
			 
			
			
			
				
					Life cycle and mating behavior					
					Maturity | Reproduction | Spawning | Eggs | Fecundity | Larvae				
				
				
				
								
				
			
			 
				
				
				
					Randall, J.E., G.R. Allen and R.C. Steene, 1990. Fishes of the Great Barrier Reef and Coral Sea. University of Hawaii Press, Honolulu, Hawaii. 506 p. (Ref. 2334)
				
				 
			
			
			
							
					
						IUCN Red List Status   (Ref. 130435: Version 2025-1)
					
					
				 
					
			
			
			
			
			
			
			
				
					Threat to humans  
				
				
				
					  Harmless				
				
			 
			
			
			
			
				
					Human uses  				
				
				
									
				
				
			
			
						
			
			
			
				
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Special reports
 
				
			
			
			
				
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					Internet sources
				
				
			
			
			
				
					Estimates based on models				
				
				
				Preferred temperature (Ref. 
123201): 26.1 - 29, mean 28 °C (based on 586 cells).
				
				
					
					Phylogenetic diversity index  (Ref. 
82804):  PD
50 = 0.5000   [Uniqueness, from 0.5 = low to 2.0 = high].					
													Bayesian length-weight: a=0.00692 (0.00314 - 0.01524), b=3.06 (2.88 - 3.24), in cm total length, based on LWR estimates for this Genus-body shape (Ref. 
93245).
					
					Trophic level  (Ref. 
69278):  3.6   ±0.4 se; based on size and trophs of closest relatives					
						 Generation time: 1.4 ( na - na) years. Estimated as median ln(3)/K based on 1
 growth studies.					
						Resilience  (Ref. 
120179):  High, minimum population doubling time less than 15 months (Preliminary K or Fecundity.).					
											
				
									
					Fishing Vulnerability  (Ref. 
59153):  Low vulnerability (18 of 100). 
🛈					
						Nutrients  (Ref. 
124155):  Calcium = 87.2 [47.6, 133.7] mg/100g; Iron = 0.685 [0.432, 1.101] mg/100g; Protein = 18.2 [15.9, 20.1] %; Omega3 = 0.0982 [, ] g/100g; Selenium = 36.4 [19.8, 69.6] μg/100g; VitaminA = 70.1 [25.2, 189.7] μg/100g; Zinc = 1.4 [1.0, 2.0] mg/100g (wet weight);