Actinopterygii (ray-finned fishes) > Perciformes
(Perch-likes) > Gobiidae
(Gobies) > Gobiinae
Etymology: Knipowitschia: Because of N.M.Knipowitsch, a zoologist and Russian ichthyologist from the Academy of Sciences. Expeditions in "Pomor" and "Pervosvanniy" ships to Azov and Black sea (1898-1908).
Environment: milieu / climate zone / depth range / distribution range
Marine; freshwater; brackish; demersal. Temperate; 4°C - 20°C (Ref. 2059); 48°N - 42°N, 30°E - 54°E
Eurasia: Brackish portions of the Black Sea, Sea of Azov and Caspian Sea.
Length at first maturity / Size / Weight / Age
Maturity: Lm 2.0  range ? - ? cm
Max length : 5.0 cm TL male/unsexed; (Ref. 2058); 3.5 cm TL (female)
(total): 6 - 8;
soft rays: 8 - 9. This species is distinguished from its congeners by the following characters: upper part of caudal fin longer than the lower part; anterior extremity of anterior oculoscapular canal in front of middle of eye; no posterior oculoscapular canal; back unscaled in front of origin of second dorsal; breeding males with 4-8 dark bars when alive; total scales in midlateral series 33-40 (Ref. 59043).
Occur in shallow water, on sand and among weeds (Ref. 4696). Live less than 2 years. Feed on small benthic invertebrates (Ref. 59043). Spawn after first winter. Females may spawn several times during a season. Males defend eggs in cavities under stones, plant material or molluscs shells (Ref. 59043) like underside of lamellibranch valves (Cardium, Monodonta, Unio, etc.) (Ref. 4696). Eggs hatch at 3.0-3.8 mm (Ref. 59043).
Males defend eggs in cavities under stones, plant material or molluscs shells (Ref. 59043). Females spawn several times during a season (Ref. 59043).
Miller, P.J., 1986. Gobiidae. p. 1019-1085. In P.J.P. Whitehead, M.-L. Bauchot, J.-C. Hureau, J. Nielsen and E. Tortonese (eds.) Fishes of the North-eastern Atlantic and the Mediterranean. Volume 3. UNESCO, Paris. (Ref. 4696)
IUCN Red List Status (Ref. 119314)
CITES (Ref. 115941)
Threat to humans
ReferencesAquacultureAquaculture profileStrainsGeneticsAllele frequenciesHeritabilityDiseasesProcessingMass conversion
Estimates of some properties based on models
Phylogenetic diversity index (Ref. 82805
= 0.5000 [Uniqueness, from 0.5 = low to 2.0 = high].
Bayesian length-weight: a=0.00891 (0.00427 - 0.01860), b=3.08 (2.91 - 3.25), in cm Total Length, based on LWR estimates for this (Sub)family-body shape (Ref. 93245
Trophic Level (Ref. 69278
): 3.4 ±0.4 se; Based on size and trophs of closest relatives
Resilience (Ref. 69278
): High, minimum population doubling time less than 15 months (tmax < 2; tm < 1 (8-10 months);).
Vulnerability (Ref. 59153
): Low vulnerability (14 of 100) .