Actinopteri (ray-finned fishes) > Gobiiformes
(Gobies) > Gobiidae
(Gobies) > Gobionellinae
Etymology: Knipowitschia: Because of N.M.Knipowitsch, a zoologist and Russian ichthyologist from the Academy of Sciences. Expeditions in "Pomor" and "Pervosvanniy" ships to Azov and Black sea (1898-1908).
Environment: milieu / climate zone / depth range / distribution range
Marine; freshwater; brackish; demersal. Temperate; 4°C - 20°C (Ref. 2059); 48°N - 42°N, 30°E - 54°E
Eurasia: Brackish portions of the Black Sea, Sea of Azov and Caspian Sea.
Length at first maturity / Size / Weight / Age
Maturity: Lm 2.0  range ? - ? cm
Max length : 5.0 cm TL male/unsexed; (Ref. 2058); 3.5 cm TL (female)
(total): 6 - 8;
soft rays: 8 - 9. This species is distinguished from its congeners by the following characters: upper part of caudal fin longer than the lower part; anterior extremity of anterior oculoscapular canal in front of middle of eye; no posterior oculoscapular canal; back unscaled in front of origin of second dorsal; breeding males with 4-8 dark bars when alive; total scales in midlateral series 33-40 (Ref. 59043).
Occur in shallow water, on sand and among weeds (Ref. 4696). Live less than 2 years. Feed on small benthic invertebrates (Ref. 59043). Spawn after first winter. Females may spawn several times during a season. Males defend eggs in cavities under stones, plant material or molluscs shells (Ref. 59043) like underside of lamellibranch valves (Cardium, Monodonta, Unio, etc.) (Ref. 4696). Eggs hatch at 3.0-3.8 mm (Ref. 59043).
Males defend eggs in cavities under stones, plant material or molluscs shells (Ref. 59043). Females spawn several times during a season (Ref. 59043).
Miller, P.J., 1986. Gobiidae. p. 1019-1085. In P.J.P. Whitehead, M.-L. Bauchot, J.-C. Hureau, J. Nielsen and E. Tortonese (eds.) Fishes of the North-eastern Atlantic and the Mediterranean. Volume 3. UNESCO, Paris. (Ref. 4696)
IUCN Red List Status (Ref. 125652)
Threat to humans
ReferencesAquacultureAquaculture profileStrainsGeneticsAllele frequenciesHeritabilityDiseasesProcessingNutrientsMass conversion
Estimates based on models
Phylogenetic diversity index (Ref. 82804
= 0.5000 [Uniqueness, from 0.5 = low to 2.0 = high].
Bayesian length-weight: a=0.00977 (0.00432 - 0.02212), b=3.09 (2.90 - 3.28), in cm total length, based on LWR estimates for this (Sub)family-body shape (Ref. 93245
Trophic level (Ref. 69278
): 3.4 ±0.4 se; based on size and trophs of closest relatives
Generation time: 1.7 ( na - na) years. Estimated as median ln(3)/K based on 2 growth studies.
Resilience (Ref. 120179
): High, minimum population doubling time less than 15 months (tmax < 2; tm < 1 (8-10 months);).
Fishing Vulnerability (Ref. 59153
): Low vulnerability (10 of 100).