Ventral margin of upper lip crenulate on lateral thirds; entire on central third; nape cirri present, simple; orbital cirri branched; 1 to 5, usually 3 to 5, preopercular pore positions with pairs or multiples of pores; subquadrate dark blotch present in humeral area on body; upper lip frequently with up to 15 alternating dark and 15 pale bands of subequal width, up to 7 of each above central, ventrally entire portion of lip; prominent, irregular dark mark on head just posterior to midpostorbital sensory pores.