a 1-3-2 interdigitation pattern; 12 caudal fin rays; 80-87 dorsal fin rays; 66-72 anal fin rays; 45-48 total vertebrae; 84-98 scales in a longitudinal series; unpigmented peritoneum; unique ocular side pigmentation featuring a yellowish to dark brown background with numerous small, dark speckles, a series of 3-8 mostly incomplete crossbands beginning just posterior to the opercle, and usually 1-3 small, dark, spherical spots arranged in a longitudinal row along the body midline; a series of dark blotches alternating with unpigmented areas on the dorsal and anal fins, and a spot and dark band on the caudal fin base with the distal four-fifths of the caudal fin unpigmented.