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|Sound produced||yes, active sound production|
|Type of sound produced||grunts, knocks|
|Sound production organ||pneumatic duct & swim bladder|
|Sound mechanism||not specified, but probably vibration of swim bladder and pneumatic duct|
|Behavioural context||when alarmed, (sound was recorded when enclosure lid was lifted)|
|Reference||Tiepelt, H., 2005|
|Remark||outdoor recording (enclosure submerged in freshwater lake). Spontaneous sound production from 1 specimen after removing the top-cover of enclosure. Sound recorded in presence of 11 others (20-25 cm long)|