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|Sound produced||yes,passive & active sound production|
|Type of sound produced||swimming noise, knocks, thumps|
|Sound production organ||swim bladder & pneumatic duct|
|Sound mechanism||may be escape sounds (discharge of gas through pneumatic duct)|
|Behavioural context||swimming & handling|
|Reference||Fish, M.P. and W.H. Mowbray, 1970|
|Remark||active sound production (knocks & thumps) only recorded during handling, recorded with bandpass filter: 30 -1200 Hz|