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|Sound produced||yes, active sound production|
|Type of sound produced||knocks, pulsed drum beats|
|Sound production organ||swim bladder|
|Sound mechanism||vibration by contraction of primary sonic muscles & conformation of swim bladder|
|Behavioural context||intraspecific competition, terretorial behaviour|
|Reference||Parmentier, E., P. Vandewalle and J.P. Lagardère, 2003|
|Remark||strongly species-specific sounds, aquarium recording, recorded with lowpass filter: 6400 Hz.|