Found in caves developed in carbonate rock of the Crawford-Mammoth Cave Uplands and Mitchell Plain in the South-Central karst region of Indiana. Its distribution is bounded to the north by the the East Fork White River and

the south by the Ohio River. Recorded from the Lower White, Lower East Fork White, Patoka and Blue-Sinking watersheds. Known from at least 74 localities

in Crawford, Harrison, Lawrence, Orange and Washington counties, including

68 cave systems and six springs. Most of these localities appear to represent sink rather source populations. Endangered in Indiana due to presumed vulnerability to groundwater pollution and other perturbations of aquatic subterranean habitats (Ref. 96073).

Status of threat: Endangered in Indiana by NatureServe (2013) due to few occurrences, small population sizes and being restricted to subterranean habitats that are highly vulnerable to anthropogenic activities (Ref. 96073).