| Main Ref. | Sokolov, L.I. and L.S. Berdicheskii, 1989 |
| Mode | dioecism |
| Fertilization | external |
| Mating type | |
| Spawning frequency | one clear seasonal peak per year |
| Spawning aggregation | Ref. |
| Batch spawner | Ref. |
| Reproductive guild |
nonguarders open water/substratum egg scatterers |
| Parental Care | none |
| Description of life cycle and mating behavior | Spawning occurs in fast-flowing water above gravel bottom, rarely among submerged vegetation. Female spawn more than once during a season, individual females spawn with several males. Males assemble at spawning site and follow ripe females, often with much splashing, to shallow riffles. Females lays sticky eggs into the gravel (Ref. 59043). Pale yellow eggs are found attached to gravel, weed and stones in flowing water (Ref. 41678). |
| Search for more references on reproduction | Scirus |