| Main Ref. | Conkel, D., 1993 | 
|---|---|
| Remarks | Inhabits lower to middle river valleys as well as lakes. Found over rocks, sand, silt or mud. Feeds on benthic detritus, aquatic plants and invertebrates. | 
| Marine - Neritic | Marine - Oceanic | Brackishwater | Freshwater | |
|---|---|---|---|---|
| Marine zones / Brackish and freshwater bodies | 
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 | 
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| Substrate | |
|---|---|
| Substrate Ref. | |
| Special habitats | |
| Special habitats Ref. | 
| Ref. | |
|---|---|
| Associations | |
| Associated with | |
| Association remarks | |
| Parasitism | 
| Feeding type | plants/detritus+animals (troph. 2.2-2.79) | 
|---|---|
| Feeding type Ref. | Conkel, D., 1993 | 
| Feeding habit | variable | 
| Feeding habit Ref. | Conkel, D., 1993 | 
| Estimation method | Original sample | Unfished population | Remark | ||
|---|---|---|---|---|---|
| Troph | s.e. | Troph | s.e. | ||
| From diet composition | |||||
| From individual food items | 2.52 | 0.20 | Trophic level estimated from a number of food items using a randomized resampling routine. | ||
| Ref. | |||||