Actinopterygii (ray-finned fishes) > Pleuronectiformes
(Flatfishes) > Bothidae
Etymology: Grammatobothus: Greek, gramma = signal, mark (Ref. 45335).
Environment: milieu / climate zone / depth range / distribution range
Marine; demersal; depth range 0 - 90 m (Ref. 9824). Tropical
Indo-West Pacific: India and off Sri Lanka eastward to the Philippines and the northeastern coast of Australia. Range extends to Okinawa Island, Japan. Also known from the Persian Gulf (Ref. 5999).
Size / Weight / Age
Maturity: Lm ?  range ? - ? cm
Max length : 21.0 cm TL male/unsexed; (Ref. 9824); common length : 17.0 cm TL male/unsexed; (Ref. 9824)
soft rays: 61 - 71. Eyed side pale brownish, with 3 large, prominent dark ocelli, 1 each above and below pectoral fin, 1 on middle of straight section of lateral line. Head and body with many small, distinct, dark spots and obscure markings along dorsal and ventral body margins. Males with 2 distinct dark markings above upper eye. Dorsal, anal, and pelvic fins on eyed side with indistinct dusky spots and markings. Caudal fin with a pair of marks behind fin base. Pectoral fin on eyed side with broad pale cross bars. Blind side pale. Body deeply ovoid, its depth 1.5 to 1.8 times in SL. Head moderately large, snout shorter than eye, profile deeply notched ahead of interorbital area. Head length3.5 to 4 times in SL. Interorbital region concave, narrow in both sexes. Both eyes on left side of head, front margin of upper eye slightly behind front margin of lower eye. Mouth moderate in size, reaching to or slightly beyond anterior margin of lower eye, length of upper jaw 2.8 to 3.4 times in head length. Teeth in both jaws uniserial. Teeth in upper jaw becoming gradually smaller posteriorly. Teeth of lower jaw about equal in size, similar to anterior teeth of upper jaw. Scales on eyed side ctenoid. Scales on blind side cycloid. Lateral line on both sides of body (Ref 42535).
Found in shallow mud, sand, and rubble bottoms of the continental shelf (Ref. 9824). Feeds on benthic animals. Marketed fresh and used in making fish meal (Ref. 9824).
Life cycle and mating behavior
Maturity | Reproduction | Spawning | Eggs | Fecundity | Larvae
Nielsen, J.G., 1974. Bothidae. In W. Fisher and P.J.P. Whitehead (eds.) FAO species identification sheets for fishery purposes. Eastern Indian Ocean (fishing area 57) and Western Central Pacific (fishing area 71). Rome, FAO, vol. 3. pag. var. (Ref. 4896)
IUCN Red List Status (Ref. 115185)
CITES (Ref. 115941)
Threat to humans
Common namesSynonymsMetabolismPredatorsEcotoxicologyReproductionMaturitySpawningFecundityEggsEgg development
ReferencesAquacultureAquaculture profileStrainsGeneticsAllele frequenciesHeritabilityDiseasesProcessingMass conversion
Estimates of some properties based on models
Preferred temperature (Ref. 115969
): 24.5 - 29, mean 28 (based on 1708 cells).
Phylogenetic diversity index (Ref. 82805
= 0.6250 [Uniqueness, from 0.5 = low to 2.0 = high].
Bayesian length-weight: a=0.01047 (0.00577 - 0.01900), b=2.94 (2.78 - 3.10), in cm Total Length, based on LWR estimates for this species & (Sub)family-body (Ref. 93245
Trophic Level (Ref. 69278
): 3.6 ±0.0 se; Based on diet studies.
Resilience (Ref. 69278
): Medium, minimum population doubling time 1.4 - 4.4 years (Preliminary K or Fecundity.).
Vulnerability (Ref. 59153
): Low to moderate vulnerability (33 of 100) .